Caesalpinia Kavaiensis
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''Mezoneuron kavaiense'' is a rare shrub or small tree in the genus '' Mezoneuron'' (pea family,
Fabaceae The Fabaceae or Leguminosae,International Code of Nomenc ...
), that is endemic to Hawaii. Common names include ''Uhiuhi'' ( the Big Island and
Kauai Kauai, () anglicized as Kauai ( ), is geologically the second-oldest of the main Hawaiian Islands (after Niʻihau). With an area of 562.3 square miles (1,456.4 km2), it is the fourth-largest of these islands and the 21st largest island ...
), ''Kāwau'' (
Maui The island of Maui (; Hawaiian: ) is the second-largest of the islands of the state of Hawaii at 727.2 square miles (1,883 km2) and is the 17th largest island in the United States. Maui is the largest of Maui County's four islands, which ...
), and ''Kea'' (Maui). It is threatened by
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
, particularly feral
ungulates Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
.


Description

''M. kavaiense'' is a
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
or small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
that reaches a height of . The bark is dark grey and made up of rectangular or oblong platelets. The
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are composed of 4 to 8 leaflets, each around in length. The
bisexual Bisexuality is a romantic or sexual attraction or behavior toward both males and females, or to more than one gender. It may also be defined to include romantic or sexual attraction to people regardless of their sex or gender identity, whi ...
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s have pink to rose
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s and red
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
. They form on pink to red terminal
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s in length. The flat, thin
seed pods This page provides a glossary of plant morphology. Botanists and other biologists who study plant morphology use a number of different terms to classify and identify plant organs and parts that can be observed using no more than a handheld magnify ...
are long, wide, and contain 2 to 4 oval-shaped
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s. Blooming takes place from December to March.


Habitat

''Uhiuhi'' inhabits dry, coastal mesic, and mixed mesic forests at elevations of . Associated plants include ''lama'' (''
Diospyros sandwicensis ''Diospyros sandwicensis'' is a species of flowering tree in the ebony family, Ebenaceae, that is endemic to Hawaii. It belongs to the same genus as both persimmons and ebony. Its common name, ''lama'', also means enlightenment in Hawaiian. ...
''), ''aalii'' (''
Dodonaea viscosa ''Dodonaea viscosa'', also known as the broadleaf hopbush, is a species of flowering plant in the ''Dodonaea'' (hopbush) genus that has a cosmopolitan distribution in Tropics, tropical, Subtropics, subtropical and warm temperate regions of Africa ...
''), and ''alahee'' (''
Psydrax odorata ''Psydrax odorata'', known as ''alahee'' in Hawaiian, is a species of flowering shrub or small tree in the coffee family, Rubiaceae. It is native to the Pacific Islands, New Guinea and Australia. Description The species range from in height, ...
''). Populations formerly existed on
Kauai Kauai, () anglicized as Kauai ( ), is geologically the second-oldest of the main Hawaiian Islands (after Niʻihau). With an area of 562.3 square miles (1,456.4 km2), it is the fourth-largest of these islands and the 21st largest island ...
(
Waimea Canyon Waimea Canyon, also known as the Grand Canyon of the Pacific, is a large canyon, approximately ten miles (16 km) long and up to 3,000 feet (900 m) deep, located on the western side of Kauai in the Hawaiian Islands of the United States. ...
),
West Maui The West Maui Mountains, West Maui Volcano, or Mauna Kahālāwai which means "holding house of water," is approximately 1.7 million years old and forms a much eroded shield volcano that constitutes the western quarter of Maui. Since its last eru ...
, Lānai, the Big Island ( North Kona District), and
Oahu Oahu () (Hawaiian language, Hawaiian: ''Oʻahu'' ()), also known as "The Gathering place#Island of Oʻahu as The Gathering Place, Gathering Place", is the third-largest of the Hawaiian Islands. It is home to roughly one million people—over t ...
(
Waianae Range Waianae () is a census-designated place (CDP) in Honolulu County, Hawaii, United States. As of the 2020 census, the CDP population was 13,614. Its name means "waters of the mullet". Its etymology is shared with the far northern Wellington subu ...
), but are only found in the latter two today.


Uses

The wood of ''M. kavaiense'' is very dense and hard, nearly black, and close-grained.
Native Hawaiians Native Hawaiians (also known as Indigenous Hawaiians, Kānaka Maoli, Aboriginal Hawaiians, First Hawaiians, or simply Hawaiians) ( haw, kānaka, , , and ), are the indigenous ethnic group of Polynesian people of the Hawaiian Islands. Hawaii ...
used it to make ''ōō'' (
digging stick A digging stick, sometimes called a yam stick, is a wooden implement used primarily by subsistence-based cultures to dig out underground food such as roots and tubers, tilling the soil, or burrowing animals and anthills. It is a term used in ar ...
s), ''ihe'' (
spear A spear is a pole weapon consisting of a shaft, usually of wood, with a pointed head. The head may be simply the sharpened end of the shaft itself, as is the case with fire hardened spears, or it may be made of a more durable material fasten ...
s), ''laau melomelo'' (
fishing lure A fishing lure is a broad type of artificial angling baits that are replicas designed to mimic real prey animals and attract the attention of predatory fish, using appearances, flashy colors, bright reflections, movements, vibrations and/or lou ...
s), ''pou'' (house posts), runners for '' papa hōlua'' (
sled A sled, skid, sledge, or sleigh is a land vehicle that slides across a surface, usually of ice or snow. It is built with either a smooth underside or a separate body supported by two or more smooth, relatively narrow, longitudinal runners ...
s), ''pāhoa'' (
dagger A dagger is a fighting knife with a very sharp point and usually two sharp edges, typically designed or capable of being used as a thrusting or stabbing weapon.State v. Martin, 633 S.W.2d 80 (Mo. 1982): This is the dictionary or popular-use de ...
s), ''laau palau'' (
clubs Club may refer to: Arts, entertainment, and media * ''Club'' (magazine) * Club, a ''Yie Ar Kung-Fu'' character * Clubs (suit), a suit of playing cards * Club music * "Club", by Kelsea Ballerini from the album ''kelsea'' Brands and enterprises ...
), and ''laau kahi wauke'' (''
Broussonetia papyrifera The paper mulberry (''Broussonetia papyrifera'', syn. ''Morus papyrifera'' L.) is a species of flowering plant in the family Moraceae. It is native to Asia, The rose-colored flowers are collected to make '' lei''. A blood purifier was made from the young leaves, leaf buds, and bark of ''uhiuhi'' mashed together with the inner bark of ''hāpuu'' (''
Cibotium ''Cibotium'' (from Greek , ''kibṓtion'', "little chest" or "box"), also known as manfern, is a genus of 11 species of tropical tree ferns. It is the only genus in family Cibotiaceae in the Pteridophyte Phylogeny Group classification of 2016 (PP ...
'' spp.), ''
okolehao Okolehao is a Hawaiian alcoholic spirit whose main ingredient was the root of the ti plant. Okolehao's forerunner was a fermented ti root beverage or beer. When distillation techniques were introduced by English seamen in 1790, it was distilled ...
'', ''ulu'' (''
Artocarpus altilis Breadfruit (''Artocarpus altilis'') is a species of flowering tree in the mulberry and jackfruit family (Moraceae) believed to be a domesticated descendant of ''Artocarpus camansi'' originating in New Guinea, the Maluku Islands, and the Philippi ...
'') bark, ''uhaloa'' (''
Waltheria indica ''Waltheria indica'' is a species of flowering plant in the mallow family, Malvaceae, that has a pantropical distribution. It is believed to have originated in the Neotropics. Common names include sleepy morning, basora prieta, hierba de soldado ...
'') taproots, and ''ko kea'' (''
Saccharum officinarum ''Saccharum officinarum'' is a large, strong-growing species of grass in the genus ''Saccharum''. Its stout stalks are rich in sucrose, a simple sugar which accumulates in the stalk internodes. It originated in New Guinea, and is now cultivated ...
'').


Conservation

The plant is very rare today, with fewer than 100 individuals remaining in the wild,The Nature Conservancy
/ref> with some estimates at fewer than 50 plants.USFWS
Determination of endangered status for Uhiuhi.
''Federal Register'' July 8, 1986.
It does not reproduce very successfully, having a low level of
recruitment Recruitment is the overall process of identifying, sourcing, screening, shortlisting, and interviewing candidates for jobs (either permanent or temporary) within an organization. Recruitment also is the processes involved in choosing individual ...
. The plant's habitat has been degraded or destroyed by development,
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to ...
, and
fire Fire is the rapid oxidation of a material (the fuel) in the exothermic chemical process of combustion, releasing heat, light, and various reaction Product (chemistry), products. At a certain point in the combustion reaction, called the ignition ...
, and invaded by
exotic species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
of plants and animals. The invasive fountaingrass (''Pennisetum setaceum'') is particularly harmful to the habitat, covering whole fields and increasing the danger of fire. The seeds are eaten by
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include ''Neotoma'' ( pack rats), ''Bandicota'' (bandicoot ...
s, and
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
and
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s graze in the area. An insect, the black coffee twig borer (''Xylosandrus compactus''), damages the
seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
s and
sapling In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s. Since the wood is so highly sought after, the trees are still in danger of being harvested. The population on the Big Island grows on the side of
Hualālai Hualālai (pronounced in Hawaiian) is an active volcano on the island of Hawaii in the Hawaiian Islands. It is the westernmost, third-youngest and the third-most active of the five volcanoes that form the island of Hawaii, following Kīlauea a ...
, a
dormant volcano A volcano is a rupture in the crust of a planetary-mass object, such as Earth, that allows hot lava, volcanic ash, and gases to escape from a magma chamber below the surface. On Earth, volcanoes are most often found where tectonic plates a ...
, and so is technically in danger of being
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
in the event of an
eruption Several types of volcanic eruptions—during which lava, tephra (ash, lapilli, volcanic bombs and volcanic blocks), and assorted gases are expelled from a volcanic vent or fissure—have been distinguished by volcanologists. These are often ...
. The plant was federally listed as an
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
of the United States in 1986.


References

* * {{Taxonbar Caesalpinieae Endemic flora of Hawaii Biota of Hawaii (island) Biota of Oahu Trees of Hawaii Plants described in 1867 Taxonomy articles created by Polbot