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The Bovidae comprise the
biological family Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of f ...
of cloven-hoofed, ruminant mammals that includes
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
,
bison Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American bison, ''B. bison'' ...
, buffalo,
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammal ...
s, and caprines. A member of this family is called a bovid. With 143
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extin ...
species and 300 known extinct species, the family Bovidae consists of 11 (or two) major subfamilies and thirteen major tribes. The family evolved 20 million years ago, in the early Miocene. The bovids show great variation in size and
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
colouration. Excepting some domesticated forms, all male bovids have two or more horns, and in many species, females possess horns, too. The size and shape of the horns vary greatly, but the basic structure is always one or more pairs of simple bony protrusions without branches, often having a spiral, twisted or fluted form, each covered in a permanent sheath of keratin. Most bovids bear 30 to 32 teeth. Most bovids are diurnal. Social activity and feeding usually peak during dawn and dusk. Bovids typically rest before dawn, during midday, and after dark. They have various methods of social organisation and
social behaviour Social behavior is behavior among two or more organisms within the same species, and encompasses any behavior in which one member affects the other. This is due to an interaction among those members. Social behavior can be seen as similar to ...
, which are classified into solitary and gregarious behaviour. Bovids use different forms of vocal, olfactory, and tangible communication. Most species alternately feed and ruminate throughout the day. While small bovids forage in dense and closed habitat, larger species feed on high-fiber vegetation in open grasslands. Most bovids are polygynous. Mature bovids mate at least once a year and smaller species may even mate twice. In some species, neonate bovids remain hidden for a week to two months, regularly nursed by their mothers; in other species, neonates are followers, accompanying their dams, rather than tending to remain hidden. The greatest diversities of bovids occur in
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. The maximum concentration of species is in the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s of Eastern Africa. Other bovid species also occur in Europe, Asia, and North America. Bovidae includes three of the five domesticated mammals whose use has spread outside their original ranges, namely cattle, sheep, and goats. Dairy products, such as milk,
butter Butter is a dairy product made from the fat and protein components of churned cream. It is a semi-solid emulsion at room temperature, consisting of approximately 80% butterfat. It is used at room temperature as a spread, melted as a condiment ...
, and
cheese Cheese is a dairy product produced in wide ranges of flavors, textures, and forms by coagulation of the milk protein casein. It comprises proteins and fat from milk, usually the milk of cows, buffalo, goats, or sheep. During production, ...
, are manufactured largely from domestic cattle. Bovids are also raised for their leather, meat, and wool.


Naming and etymology

The name "Bovidae" was given by the British zoologist John Edward Gray in 1821. The word "Bovidae" is the combination of the prefix ''bov-'' (originating from Latin ''bos'', "ox", through Late Latin ''bovinus'') and the suffix ''-idae''.


Taxonomy

The family Bovidae is placed in the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
Artiodactyla The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
(which includes the even-toed
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, c ...
s). It includes 143 extant species, accounting for nearly 55% of the
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, c ...
s, and 300 known extinct species. Until the beginning of the 21st century it was understood that the family
Moschidae Moschidae is a family of pecoran even-toed ungulates, containing the musk deer (''Moschus'') and its extinct relatives. They are characterized by long 'saber teeth' instead of horns, antlers or ossicones, modest size (''Moschus'' only reaches ; o ...
(musk deer) was sister to
Cervidae Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
. However, a 2003
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study by Alexandre Hassanin (of National Museum of Natural History, France) and colleagues, based on mitochondrial and
nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: *Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics *Nuclear space *Nuclear ...
analyses, revealed that
Moschidae Moschidae is a family of pecoran even-toed ungulates, containing the musk deer (''Moschus'') and its extinct relatives. They are characterized by long 'saber teeth' instead of horns, antlers or ossicones, modest size (''Moschus'' only reaches ; o ...
and Bovidae form a clade sister to
Cervidae Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
. According to the study, Cervidae diverged from the Bovidae-Moschidae clade 27 to 28 million years ago. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
is based on the 2003 study. Molecular studies have supported monophyly in the family Bovidae (a group of organisms comprises an ancestral species and all their descendants). The number of subfamilies in Bovidae is disputed, with suggestions of as many as ten and as few as two subfamilies. However, molecular, morphological and fossil evidence indicates the existence of eight distinct subfamilies: Aepycerotinae (consisting of just the impala),
Alcelaphinae The subfamily Alcelaphinae or tribe Alcelaphini of the family Bovidae contains wildebeest, hartebeest, bonteboks, and several similar species. Depending on the classification, there are 6–10 species placed in four genera, although ''Beatragu ...
(bontebok, hartebeest, wildebeest and relatives),
Antilopinae The Antilopines are even-toed ungulates belonging to the subfamily Antilopinae of the family Bovidae. The members of tribe Antilopini include the gazelles, blackbucks, springboks, gerenuks, dibatags, and Central Asian gazelles, are often referred ...
(several antelopes, gazelles, and relatives),
Bovinae Bovines (subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship betwe ...
(cattle, buffaloes, bison and other antelopes),
Caprinae The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine, or, more informally, a goat-antelope ( ...
(goats, sheep, ibex, serows and relatives), Cephalophinae (duikers), Hippotraginae (addax, oryx and relatives) and
Reduncinae The bovid subfamily Reduncinae or tribe Reduncini is composed of nine species of antelope, all of which dwell in marshes, floodplains, or other well-watered areas, including the waterbucks and reedbucks. These antelopes first appear in the fossil ...
(reedbuck and kob antelopes). In addition, three extinct subfamilies are known:
Hypsodontinae Hypsodontinae is an extinct subfamily of Bovidae. They are considered the earliest members of the family, first appearing in the earliest part of the Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from ...
(mid- Miocene), Oiocerinae (
Turolian The Turolian age is a period of geologic time The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pr ...
) and the subfamily Tethytraginae, which contains '' Tethytragus'' (mid- Miocene). In 1992, Alan W. Gentry of the Natural History Museum, London divided the eight major subfamilies of Bovidae into two major clades on the basis of their evolutionary history: the Boodontia, which comprised only the Bovinae, and the Aegodontia, which consisted of the rest of the subfamilies. Boodonts have somewhat primitive teeth, resembling those of oxen, whereas aegodonts have more advanced teeth like those of goats. A controversy exists about the recognition of
Peleinae The grey rhebok or gray rhebuck (''Pelea capreolus''), locally known as the ''reebok'' in Afrikaans, is a species of antelope native to South Africa, Lesotho, and Eswatini (Swaziland). The specific name ''capreolus'' is Latin for 'little goat' ...
and Pantholopinae, comprising the genera '' Pelea'' and '' Pantholops'' respectively, as subfamilies. In 2000, American biologist George Schaller and palaeontologist Elisabeth Vrba suggested the inclusion of '' Pelea'' in
Reduncinae The bovid subfamily Reduncinae or tribe Reduncini is composed of nine species of antelope, all of which dwell in marshes, floodplains, or other well-watered areas, including the waterbucks and reedbucks. These antelopes first appear in the fossil ...
, though the grey rhebok, the sole species of '' Pelea'', is highly different from kobs and reduncines in morphology. '' Pantholops'', earlier classified in the
Antilopinae The Antilopines are even-toed ungulates belonging to the subfamily Antilopinae of the family Bovidae. The members of tribe Antilopini include the gazelles, blackbucks, springboks, gerenuks, dibatags, and Central Asian gazelles, are often referred ...
, was later placed in its own subfamily, Pantholopinae. However, molecular and morphological analysis supports the inclusion of '' Pantholops'' in
Caprinae The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine, or, more informally, a goat-antelope ( ...
. Below is a cladogram based on Yang ''et al''.'','' 2013 and Calamari, 2021: Alternatively, all members of the Aegodontia, can be classified within the subfamily Antilopinae, with the individual subfamilies being tribes in this treatment.


Evolution


Early Miocene and before

In the early Miocene, bovids began diverging from the
cervid Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
s (deer) and giraffids. The earliest bovids, whose presence in Africa and Eurasia in the latter part of early Miocene (20 Mya) has been ascertained, were small animals, somewhat similar to modern gazelles, and probably lived in woodland environments. ''
Eotragus ''Eotragus'' is an extinct genus of early bovid. Species belonging to the genus inhabited Europe, Africa, and Asia during the Miocene some 20-18 million years ago. It is related to the modern nilgai and four-horned antelope. It was small and ...
'', the earliest known bovid, weighed and was nearly the same in size as the Thomson's gazelle. Early in their evolutionary history, the bovids split into two main clades: Boodontia (of Eurasian origin) and Aegodontia (of African origin). This early
split Split(s) or The Split may refer to: Places * Split, Croatia, the largest coastal city in Croatia * Split Island, Canada, an island in the Hudson Bay * Split Island, Falkland Islands * Split Island, Fiji, better known as Hạfliua Arts, enterta ...
between Boodontia and Aegodontia has been attributed to the continental divide between these land masses. When these continents were later rejoined, this barrier was removed, and either group expanded into the territory of the other. The tribes
Bovini The tribe Bovini, or wild cattle, are medium to massive bovines that are native to North America, Eurasia, and Africa. These include the enigmatic, antelope-like saola, the African and Asiatic buffalos, and a clade that consists of bison and the ...
and Tragelaphini diverged in the early Miocene. Bovids are known to have reached the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with t ...
in the Pleistocene by crossing the
Bering land bridge Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip o ...
. The present genera of Alcelaphinae appeared in the Pliocene. The extinct Alcelaphine genus ''Paramularius'', which was the same in size as the hartebeest, is believed to have come into being in the Pliocene, but became extinct in the middle Pleistocene. Several genera of Hippotraginae are known since the Pliocene and Pleistocene. This subfamily appears to have diverged from the Alcelaphinae in the latter part of early Miocene. The Bovinae are believed to have diverged from the rest of the Bovidae in the early Miocene. The Boselaphini became extinct in Africa in the early Pliocene; their latest fossils were excavated in
Langebaanweg Langebaanweg is a town on the southwest coast of South Africa, in Western Cape Province. It is the location of the air force base AFB Langebaanweg. Langebaanweg has been an important mining center, with its mai minerals consisting of phosphorit ...
(South Africa) and
Lothagam Lothagam is a geological formation located in Kenya, near the southwestern shores of Lake Turkana, from Kanapoi. It is located between the Kerio and Lomunyenkuparet Rivers on an uplifted fault block. Lothagam has deposits dating to the Miocene-P ...
(Kenya).


Middle Miocene

The middle Miocene marked the spread of the bovids into China and the Indian subcontinent. According to Vrba, the radiation of the subfamily Alcelaphinae began in the latter part of middle Miocene. The Caprinae tribes probably diverged in the early middle Miocene. The Caprini emerged in the middle Miocene, and seem to have been replaced by other bovids and cervids in Eurasia. The earliest fossils of the antilopines are from the middle Miocene, though studies show the existence of the subfamily from the early Miocene.
Speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
occurred in the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to conflic ...
Antilopini Antilopini is a tribe of bovids often referred as true antelopes like gazelles. They live in and around the Sahara, Horn of Africa, throughout eastern and southern Africa, and Eurasia. Depending on species, the females have either very short and/ ...
during the middle or upper Miocene, mainly in Eurasia. Tribe
Neotragini The tribe Neotragini comprises the dwarf antelopes of Africa: * '' Dorcatragus'' ** Beira ''D. megalotis'' * '' Madoqua'' ** Günther's dik-dik ''M. guentheri'' ** Kirk's dik-dik ''M. kirkii'' ** Silver dik-dik ''M. piacentinii'' ** Salt's d ...
seems to have appeared in Africa by the end of Miocene, and had become widespread by the Pliocene.


Late Miocene

By the late Miocene, around 10 Mya, the bovids rapidly diversified, leading to the creation of 70 new genera. This late Miocene radiation was partly because many bovids became adapted to more open, grassland habitats. The Aepycerotinae first appeared in the late Miocene, and no significant difference in the sizes of the primitive and modern impala has been noted. Fossils of ovibovines, a tribe of Caprinae, in Africa date back to the late Miocene. The earliest Hippotragine fossils date back to the late Miocene, and were excavated from sites such as Lothagam and
Awash Valley The Awash (sometimes spelled Awaash; Oromo: ''Awaash'', Amharic: አዋሽ, Afar: ''We'ayot'', Somali: ''Webiga Dir'') is a major river of Ethiopia. Its course is entirely contained within the boundaries of Ethiopia and empties into a chain of ...
. The first African fossils of Reduncinae date back to 6-7 Mya. Reduncinae and Peleinae probably diverged in the mid-Miocene.


Characteristics

All bovids have the similar basic form - a snout with a blunt end, one or more pairs of horns (generally present on males) immediately after the oval or pointed ears, a distinct neck and limbs, and a tail varying in length and bushiness among the species. Most bovids exhibit sexual dimorphism, with males usually larger as well as heavier than females. Sexual dimorphism is more prominent in medium- to large-sized bovids. All bovids have four toes on each foot – they walk on the central two (the hooves), while the outer two (the
dewclaw A dewclaw is a digit – vestigial in some animals – on the foot of many mammals, birds, and reptiles (including some extinct orders, like certain theropods). It commonly grows higher on the leg than the rest of the foot, such that in digi ...
s) are much smaller and rarely touch the ground. The bovids show great variation in size: the gaur can weigh more than , and stand high at the shoulder. The
royal antelope The royal antelope (''Neotragus pygmaeus'') is a West African antelope, recognized as the world's smallest antelope. It was first described by Swedish zoologist Carl Linnaeus in 1758. It stands up to merely at the shoulder and weighs . A chara ...
, in sharp contrast, is only tall and weighs at most . The klipspringer, another small antelope, stands at the shoulder and weighs just . Differences occur in
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
colouration, ranging from a pale white (as in the
Arabian oryx The Arabian oryx (''Oryx leucoryx'') or white oryx is a medium-sized antelope with a distinct shoulder bump, long, straight horns, and a tufted tail. It is a bovid, and the smallest member of the genus ''Oryx'', native to desert and steppe areas o ...
) to black (as in the black wildebeest). However, only the intermediate shades, such as brown and reddish brown (as in the
reedbuck Reedbuck is a common name for Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it ...
), are commonly observed. In several species, females and juveniles exhibit a light-coloured coat, while those of males darken with age. As in the
wildebeest Wildebeest ( , , ), also called gnu ( or ), are antelopes of the genus ''Connochaetes'' and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toe ...
, the coat may be marked with prominent or faint stripes. In some species such as the
addax The addax (''Addax nasomaculatus''), also known as the white antelope and the screwhorn antelope, is an antelope native to the Sahara Desert. The only member of the genus ''Addax'', it was first described scientifically by Henri de Blainvil ...
, the coat colour can vary by the season. Scent glands and sebaceous glands are often present. Some species, such as the gemsbok, sable antelope, and
Grant's gazelle Grant's gazelle (''Nanger granti'') is a species of gazelle distributed from northern Tanzania to South Sudan and Ethiopia, and from the Kenyan coast to Lake Victoria. Its Swahili name is ''swala granti''.camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
d with strongly disruptive facial markings that conceal the highly recognisable eye. Many species, such as gazelles, may be made to look flat, and hence to blend into the background, by
countershading Countershading, or Thayer's law, is a method of camouflage in which an animal's coloration is darker on the top or upper side and lighter on the underside of the body. This pattern is found in many species of mammals, reptiles, birds, fish, and ...
. The outlines of many bovids are broken up with bold disruptive colouration, the strongly contrasting patterns helping to delay recognition by predators. However, all the Hippotraginae (including the gemsbok) have pale bodies and faces with conspicuous markings. The zoologist
Tim Caro Timothy M. Caro (born 1951) is a British evolutionary ecologist known for his work on conservation biology, animal behaviour, anti-predator defences in animals, and the function of zebra stripes. He is the author of several textbooks on these subj ...
describes this as difficult to explain, but given that the species are diurnal, he suggests that the markings may function in communication. Strongly contrasting leg colouration is common only in the Bovidae, where for example ''Bos'', ''Ovis'', bontebok and gemsbok have white stockings. Again, communication is the likely function. Excepting some domesticated forms, all male bovids have horns, and in many species, females, too, possess horns. The size and shape of the horns vary greatly, but the basic structure is a pair of simple bony protrusions without branches, often having a spiral, twisted, or fluted form, each covered in a permanent sheath of keratin. Although horns occur in a single pair on almost all bovid species, there are exceptions such as the
four-horned antelope The four-horned antelope (''Tetracerus quadricornis''), or ''chousingha'', is a small antelope found in India and Nepal. Its four horns distinguish it from most other bovids, which have two horns (with a few exceptions, such as the Jacob sheep) ...
and the Jacob sheep. The unique horn structure is the only unambiguous morphological feature of bovids that distinguishes them from other
pecorans Pecora is an infraorder of even-toed hoofed mammals with ruminant digestion. Most members of Pecora have cranial appendages projecting from their frontal bones; only two extant genera lack them, '' Hydropotes'' and ''Moschus''. The name “Pec ...
. A high correlation exists between horn morphology and fighting behaviour of the individual. For instance, long horns are intended for wrestling and fencing, whereas curved horns are used in ramming. Males with horns directed inwards are monogamous and solitary, while those with horns directed outwards tend to be polygynous. These results were independent of body size. Male horn development has been linked to
sexual selection Sexual selection is a mode of natural selection in which members of one biological sex choose mates of the other sex to mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex (int ...
, Horns are small spikes in the monogamous duikers and other small antelopes, whereas in the polygynous, they are large and elaborately formed (for example in a spiral structure, as in the
giant eland The giant eland (''Taurotragus derbianus''), also known as the Lord Derby eland and greater eland, is an open-forest and savanna antelope. A species of the family Bovidae and genus ''Taurotragus'', it was described in 1847 by John Edward Gray. T ...
). Thus, to some extent, horns depict the degree of
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indivi ...
among males in a species. However, the presence of horns in females is likely due to natural selection. The horns of females are usually smaller than those of males, and are sometimes of a different shape. The horns of female bovids are believed to have evolved for defence against predators or to express territoriality, as nonterritorial females, which are able to use
crypsis In ecology, crypsis is the ability of an animal or a plant to avoid observation or detection by other animals. It may be a predation strategy or an antipredator adaptation. Methods include camouflage, nocturnality, subterranean lifestyle and m ...
for predator defence, often do not have horns. Females possess horns only in half of the bovid genera, and females in these genera are heavier than those in the rest. Females use horns mainly for stabbing.


Anatomy

In bovids, the third and fourth
metapodial Metapodials are long bones of the hand (metacarpals) and feet (metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes ...
s are combined into the
cannon bone Good conformation in the limbs leads to improved movement and decreased likelihood of injuries. Large differences in bone structure and size can be found in horses used for different activities, but correct conformation remains relatively simil ...
. The
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
and fibula are reduced, and fused with the radius and tibia, respectively. Long
scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...
e are present, whereas the clavicles are absent. Being
ruminants Ruminants (suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
, the stomach is composed of four chambers: the rumen (80%), the
omasum The omasum, also known as the bible, the fardel, the manyplies and the psalterium, is the third compartment of the stomach in ruminants. The omasum comes after the rumen and reticulum and before the abomasum. Different ruminants have different om ...
, the
reticulum Reticulum is a small, faint constellation in the southern sky. Its name is Latin for a small net, or reticle—a net of crosshairs at the focus of a telescope eyepiece that is used to measure star positions. The constellation is best viewed be ...
, and the
abomasum The abomasum, also known as the maw,The Cham ...
. The
ciliate The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to eukaryotic flagella, but are in general shorter and present in much larger numbers, with a differen ...
s and bacteria of the rumen
ferment Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food p ...
the complex
cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
into simpler fatty acids, which are then absorbed through the rumen wall. Bovids have a long
small intestine The small intestine or small bowel is an organ in the gastrointestinal tract where most of the absorption of nutrients from food takes place. It lies between the stomach and large intestine, and receives bile and pancreatic juice through the ...
; the length of the small intestine in
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult ma ...
is . Body temperature fluctuates through the day; for instance, in goats the temperature can change slightly from nearly in the early morning to in the afternoon. Temperature is regulated through sweating in cattle, whereas goats use panting for the same. The right lung, consisting of four to five lobes, is around 1.5 times larger than the left, which has three lobes.


Dentition

Most bovids bear 30 to 32 teeth. While the upper incisors are absent, the upper
canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the sur ...
are either reduced or absent. Instead of the upper incisors, bovids have a thick and tough layer of tissue, called the
dental pad The dental pad or browsing pad is a feature of ruminant dental anatomy that results from a lack of upper incisors and helps them gather large quantities of grass and other plant matter. This feature can be found in ruminants such as cattle and she ...
, that provides a surface to grip grasses and foliage. They are
hypsodont Hypsodont is a pattern of dentition with high-crowned teeth and enamel extending past the gum line, providing extra material for wear and tear. Some examples of animals with hypsodont dentition are cows and horses; all animals that feed on gritty ...
and selenodont, since the molars and
premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mou ...
s are low-
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
ed and crescent-shaped cusps. The lower incisors and canines project forward. The incisors are followed by a long toothless gap, known as the diastema. The general dental formula for bovids is . Most members of the family are herbivorous, but most duikers are
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
. Like other ruminants, bovids have four-chambered stomachs, which allow them to digest plant material, such as
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
, that cannot be used by many other animals. Ruminants (and some others like kangaroos, rabbits, and
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s) are able to use
micro-organisms A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
living in their guts to break down cellulose by fermentation.


Ecology and behaviour

The bovids have various methods of social organisation and social behaviour, which are classified into solitary and gregarious behaviour. Further, these types may each be divided into territorial and nonterritorial behaviour. Small bovids such as the klipspringer,
oribi The oribi (; ''Ourebia ourebi'') is a small antelope found in eastern, southern and western Africa. The sole member of its genus, it was described by the German zoologist Eberhard August Wilhelm von Zimmermann in 1783. While this is the only m ...
, and
steenbok The steenbok (''Raphicerus campestris'') is a common small antelope of southern and eastern Africa. It is sometimes known as the steinbuck or steinbok. Description Steenbok resemble small oribi, standing 45–60 cm (16"–24") at the s ...
are generally solitary and territorial. They hold small territories into which other members of the species are not allowed to enter. These antelopes form monogamous pairs. Many species such as the dik-dik use pheromone secretions from the
preorbital gland The preorbital gland is a paired exocrine gland found in many species of hoofed animals, which is homologous to the lacrimal gland found in humans. These glands are trenchlike slits of dark blue to black, nearly bare skin extending from the med ...
s and sometimes dung, as well, to mark their territories. The offspring disperse at the time of adolescence, and males must acquire territories prior to mating. The bushbuck is the only bovid that is both solitary and not territorial. This antelope hardly displays aggression, and tends to isolate itself or form loose herds, though in a favourable habitat, several bushbuck may be found quite close to one another. Excluding the cephalophines (duikers), tragelaphines (spiral-horned antelopes) and the neotragines, most African bovids are gregarious and territorial. Males are forced to disperse on attaining sexual maturity, and must form their own territories, while females are not required to do so. Males that do not hold territories form bachelor herds. Competition takes place among males to acquire dominance, and fights tend to be more rigorous in limited
rutting season The rut (from the Latin ''rugire'', meaning "to roar") is the mating season of certain mammals, which includes ruminants such as deer, sheep, camels, goats, pronghorns, bison, giraffes and antelopes, and extends to others such as skunks and eleph ...
s. With the exception of migratory males, males generally hold the same territory throughout their lives. In the waterbuck, some male individuals, known as "satellite males", may be allowed into the territories of other males and have to wait till the owner grows old so they may acquire his territory.
Lek mating A lek is an aggregation of male animals gathered to engage in competitive displays and courtship rituals, known as lekking, to entice visiting females which are surveying prospective partners with which to mate. A lek can also indicate an avail ...
, where males gather together and competitively display to potential mates, is known to exist among topis,
kob The kob (''Kobus kob'') is an antelope found across Central Africa and parts of West Africa and East Africa. Together with the closely related reedbucks, waterbucks, lechwe, Nile lechwe, and puku, it forms the Reduncinae tribe. Found along t ...
s, and lechwes. The tragelaphines, cattle, sheep, and goats are gregarious and not territorial. In these species, males must gain absolute dominance over all other males, and fights are not confined to territories. Males, therefore, spend years in body growth.


Activity

Most bovids are diurnal, although a few such as the buffalo, bushbuck, reedbuck, and grysbok are exceptions. Social activity and feeding usually peak during dawn and dusk. The bovids usually rest before dawn, during midday, and after dark. Grooming is usually by licking with the tongue. Rarely do antelopes roll in mud or dust. Wildebeest and buffalo usually wallow in mud, whereas the hartebeest and topi rub their heads and horns in mud and then smear it over their bodies. Bovids use different forms of vocal, olfactory, and tangible communication. These involve varied postures of neck, head, horns, hair, legs, and ears to convey sexual excitement, emotional state, or alarm. One such expression is the
flehmen response The flehmen response (; from German ''flehmen'', to bare the upper teeth, and Upper Saxon German ''flemmen'', to look spiteful), also called the flehmen position, flehmen reaction, flehmen grimace, flehming, or flehmening, is a behavior in whic ...
. Bovids usually stand motionless, with the head high and an intent stare, when they sense danger. Some like the impala, kudu, and eland can even leap to heights of a few feet. Bovids may roar or grunt to caution others and warn off predators. Bovids such as gazelles stot or pronk in response to predators, making high leaps on stiff legs, indicating honestly both that the predator has been seen, and that the stotting individual is strong and not worth chasing. In the mating season, rutting males bellow to make their presence known to females. Muskoxen roar during male-male fights, and male saigas force air through their noses, producing a roar to deter rival males and attract females. Mothers also use vocal communication to locate their calves if they get separated. During fights over dominance, males tend to display themselves in an erect posture with a level muzzle. Fighting techniques differ amongst the bovid families and also depend on their build. While the hartebeest fight on knees, others usually fight on all fours. Gazelles of various sizes use different methods of combat. Gazelles usually box, and in serious fights may clash and fence, consisting of hard blows from short range. Ibex, goat and sheep males stand upright and clash into each other downwards. Wildebeest use powerful head butting in aggressive clashes. If horns become entangled, the opponents move in a circular manner to unlock them. Muskoxen will ram into each other at high speeds. As a rule, only two bovids of equal build and level of defence engage in a fight, which is intended to determine the superior of the two. Individuals that are evidently inferior to others would rather flee than fight; for example, immature males do not fight with the mature bulls. Generally, bovids direct their attacks on the opponent's head rather than its body. The S-shaped horns, such as those on the impala, have various sections that help in ramming, holding, and stabbing. Serious fights leading to injury are rare.


Diet

Most bovids alternately feed and ruminate throughout the day. While those that feed on concentrate feed and digest in short intervals, the roughage feeders take longer intervals. Only small species such as the duiker browse for a few hours during day or night. Feeding habits are related to body size; while small bovids forage in dense and closed habitat, larger species feed upon high-fiber vegetation in open grasslands. Subfamilies exhibit different feeding strategies. While Bovinae species graze extensively on fresh grass and diffused forage, Cephalophinae species (with the exception of ''
Sylvicapra The common duiker (''Sylvicapra grimmia''), also known as the grey or bush duiker, is a small antelope and the only member of the genus ''Sylvicapra''. This species is found everywhere in Africa south of the Sahara, excluding the Horn of Africa ...
'') primarily consume fruits. Reduncinae and Hippotraginae species depend on unstable food sources, but the latter are specially adapted to arid areas. Members of Caprinae, being flexible feeders, forage even in areas with low productivity. Tribes Alcelaphini, Hippotragini, and Reduncini have high proportions of monocots in their diets. On the contrary, Tragelaphini and Neotragini (with the exception of ''
Ourebia The oribi (; ''Ourebia ourebi'') is a small antelope found in eastern, southern and western Africa. The sole member of its genus, it was described by the German zoologist Eberhard August Wilhelm von Zimmermann in 1783. While this is the only m ...
'') feed extensively on dicots. No conspicuous relationship exists between body size and consumption of monocots.


Sexuality and reproduction

Most bovids are polygynous. In a few species, individuals are monogamous, resulting in minimal male-male aggression and reduced selection for large body size in males. Thus, sexual dimorphism is almost absent. Females may be slightly larger than males, possibly due to competition among females for the acquisition of territories. This is the case in duikers and other small bovids. The time taken for the attainment of
sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definitio ...
by either sex varies broadly among bovids. Sexual maturity may even precede or follow mating. For instance, the impala males, though sexually mature by a year, can mate only after four years of age. On the contrary
barbary sheep The Barbary sheep (''Ammotragus lervia''), also known as aoudad (pronounced �ɑʊdæd is a species of caprine native to rocky mountains in North Africa. While this is the only species in genus ''Ammotragus'', six subspecies have been describe ...
females may give birth to offspring even before they have gained sexual maturity. The delay in male sexual maturation is more visible in sexually dimorphic species, particularly the reduncines, probably due to competition among males. For instance, the blue wildebeest females become capable of reproduction within a year or two of birth, while the males become mature only when four years old. All bovids mate at least once a year, and smaller species may even mate twice. Mating seasons occur typically during the rainy months for most bovids. As such, breeding might peak twice in the equatorial regions. The sheep and goats exhibit remarkable seasonality of reproduction, in the determination of which the annual cycle of daily
photoperiod Photoperiodism is the physiological reaction of organisms to the length of night or a dark period. It occurs in plants and animals. Plant photoperiodism can also be defined as the developmental responses of plants to the relative lengths of light a ...
plays a pivotal role. Other factors that have a significant influence on this cycle include the temperature of the surroundings, nutritional status, social interactions, the date of parturition and the lactation period. A study of this phenomenon concluded that goats and sheep are short-day breeders. Mating in most sheep breeds begins in summer or early autumn. Mating in sheep is also affected by melatonin, that advances the onset of the breeding season; and thyroxine, that terminates the breeding season.
Estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
lasts for at most a day in bovids, with the exception of bovines and tragelaphines. Except for the hartebeest and the topi, all bovids can detect estrus in females by testing the urine using the vomeronasal organ. Once the male is assured that the female is in estrus, he begins courtship displays; these displays vary greatly from the elaborate marches among gregarious species to the fervent licking of female genitalia among solitary species. Females, initially not receptive, ultimately mates with the male which has achieved dominance over others. Receptiveness is expressed by permission for mounting by the male and setting aside the tail by the female. Copulation generally takes a few seconds. Gestational period varies among bovids - while duiker gestation ranges from 120 to 150 days, gestation in African buffalo ranges from 300 to 330 days. Usually, a single offspring is born (twins are less frequent), and it is able to stand and run by itself within an hour of birth. In monogamous species, males assist in defending their young, but that is not the case in polygynous species. Most newborn calves remain hidden for a week to two months, regularly nursed by their mothers. In some bovid species, the neonates start following about their mothers immediately or within a few days, as in the impala. Different bovids have different strategies for the defence of juveniles. For instance, while wildebeest mothers solely defend their young, buffaloes exhibit collective defence. Weaning might occur as early as two months (as in royal antelope) or as late as a year (as in
muskox The muskox (''Ovibos moschatus'', in Latin "musky sheep-ox"), also spelled musk ox and musk-ox, plural muskoxen or musk oxen (in iu, ᐅᒥᖕᒪᒃ, umingmak; in Woods Cree: ), is a hoofed mammal of the family Bovidae. Native to the Arctic, ...
).


Lifespan

Most wild bovids live for 10 to 15 years. Larger species tend to live longer; for instance,
American bison The American bison (''Bison bison'') is a species of bison native to North America. Sometimes colloquially referred to as American buffalo or simply buffalo (a different clade of bovine), it is one of two extant species of bison, alongside the ...
can live up to 25 years and gaur up to 30 years. The mean lifespan of domesticated individuals is nearly ten years. For example, domesticated goats have an average lifespan of 12 years. Usually males, mainly in polygynous species, have shorter lifespans than females. This can be attributed to several reasons: early dispersal of young males, aggressive male-male fights, vulnerability to predation (particularly when males are less agile, as in kudu), and malnutrition (being large in size, the male body has high nutritional requirements which may not be satisfied). Richard Despard Estes suggested that females mimic male secondary sexual characteristics like horns to protect their male offspring from dominant males. This feature seems to have been strongly selected to prevent male mortality and imbalanced sex ratios due to attacks by aggressive males and forced dispersal of young males during
adolescence Adolescence () is a transitional stage of physical and psychological development that generally occurs during the period from puberty to adulthood (typically corresponding to the age of majority). Adolescence is usually associated with t ...
.


Distribution

Most of the diverse bovid species occur in Africa. The maximum concentration is in the
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s of eastern Africa. Depending on their feeding habits, several species have radiated over large stretches of land, and hence several variations in dental and limb morphology are observed. Duikers inhabit the equatorial rainforests,
sitatunga The sitatunga or marshbuck (''Tragelaphus spekii'') is a swamp-dwelling antelope found throughout central Africa, centering on the Democratic Republic of the Congo, the Republic of the Congo, Cameroon, parts of Southern Sudan, Equatorial Guinea ...
, and lechwe occur near swamps, eland inhabit grasslands, springbok and oryx occur in deserts, bongo and
anoa Anoa, also known as dwarf buffalo and ''sapiutan'', are two species of the genus ''Bubalus'' endemic to the island of Sulawesi in Indonesia: the mountain anoa (''Bubalus quarlesi'') and the lowland anoa (''Bubalus depressicornis''). Both live ...
live in dense forests, and mountain goats and takin live at high altitudes. A few bovid species also occur in Europe, Asia, and North America. Sheep and goats are found primarily in Eurasia, though the Barbary sheep and the ibex form part of the African fauna. The muskox is confined to the
arctic tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
. Several bovid species have been domesticated by human beings. The domestication of goats and sheep began 10 thousand years ago, while cattle were domesticated about 7.5 thousand years ago.


Interaction with humans


Domesticated animals

The domestication of bovids has contributed to shifting the dependence of human beings from hunting and gathering to agriculture. The Bovidae includes three of the six large domesticated herbivores whose use has spread outside their original ranges, namely cattle, sheep, and goats; all are from Eurasia, and are now found across the world. The other three species are the horse, donkey, and pig. Other large bovids that have been domesticated but which remain within the ranges of their wild ancestors are the water buffalo (from the Indian water buffalo), domestic yak (from the
wild yak The wild yak (''Bos mutus'') is a large, wild bovine native to the Himalayas. It is the ancestor of the domestic yak (''Bos grunniens''). Taxonomy The ancestor of the wild and domestic yak is thought to have diverged from ''Bos primigenius'' ...
), zebu (from the Indian aurochs), gayal (from the gaur) and
Bali cattle The Bali cattle (''Bos javanicus domesticus''), also known as Balinese cattle, domestic banteng or Bali banteng, are a domesticated species of bovine which originated from the banteng (''Bos javanicus''). Bali cattle are an important source of ...
(from the
banteng The banteng (''Bos javanicus''; ), also known as tembadau, is a species of cattle found in Southeast Asia. The head-and-body length is between . Wild banteng are typically larger and heavier than their domesticated counterparts, but are otherw ...
). Some antelopes have been domesticated including the
oryx ''Oryx'' is a genus consisting of four large antelope species called oryxes. Their pelage is pale with contrasting dark markings in the face and on the legs, and their long horns are almost straight. The exception is the scimitar oryx, which ...
es,
addax The addax (''Addax nasomaculatus''), also known as the white antelope and the screwhorn antelope, is an antelope native to the Sahara Desert. The only member of the genus ''Addax'', it was first described scientifically by Henri de Blainvil ...
, elands and the extinct
bubal hartebeest The bubal hartebeest, also known as northern hartebeest or bubal antelope or simply bubal (''Alcelaphus buselaphus buselaphus'') is the extinct nominal (i.e., first described) subspecies of hartebeest. It was formerly found north of the Saharan ...
. In Ancient Egypt oryxes, addaxes and bubal hartebeests are depicted in carved walls. The earliest evidence of cattle domestication is from 8000 BC, suggesting that the process began in Cyprus and the Euphrates basin.


Animal products

Dairy products such as milk,
butter Butter is a dairy product made from the fat and protein components of churned cream. It is a semi-solid emulsion at room temperature, consisting of approximately 80% butterfat. It is used at room temperature as a spread, melted as a condiment ...
, ghee,
yoghurt Yogurt (; , from tr, yoğurt, also spelled yoghurt, yogourt or yoghourt) is a food produced by bacterial fermentation of milk. The bacteria used to make yogurt are known as ''yogurt cultures''. Fermentation of sugars in the milk by these bact ...
,
buttermilk Buttermilk is a fermented dairy drink. Traditionally, it was the liquid left behind after churning butter out of cultured cream. As most modern butter in western countries is not made with cultured cream but uncultured sweet cream, most mod ...
and
cheese Cheese is a dairy product produced in wide ranges of flavors, textures, and forms by coagulation of the milk protein casein. It comprises proteins and fat from milk, usually the milk of cows, buffalo, goats, or sheep. During production, ...
are manufactured largely from domestic cattle, though the milk of sheep, goat, yak, and buffalo is also used in some parts of the world and for gourmet products. For example, buffalo milk is used to make
mozzarella Mozzarella (, ; nap, muzzarella ) is a southern Italian cheese traditionally made from Italian buffalo's milk by the pasta filata method. Fresh mozzarella is generally white but when seasoned it turns to a light yellow depending on the anim ...
in Italy and '' gulab jamun'' dessert in India, while sheep milk is used to make blue
Roquefort Roquefort is a sheep milk cheese from Southern France, and is one of the world's best known blue cheeses. Though similar cheeses are produced elsewhere, EU law dictates that only those cheeses aged in the natural Combalou caves of Roquefort-sur ...
cheese in France.
Beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantity ...
is a food source high in zinc, selenium, phosphorus, iron, and
B vitamin B vitamins are a class of water-soluble vitamins that play important roles in cell metabolism and synthesis of red blood cells. Though these vitamins share similar names (B1, B2, B3, etc.), they are chemically distinct compounds that often coexis ...
s. Bison meat is lower in fat and cholesterol than beef, but has a higher protein content. Bovid leather is tough and durable, with the additional advantage that it can be made into leathers of varying thicknesses - from soft clothing leather to hard shoe leather. While goat and cattle leather have a wide variety of use, sheepskin is suited only for clothing purposes. Wool from Merino hoggets is the finest and most valuable.
Merino wool The Merino is a breed or group of breeds of domestic sheep, characterised by very fine soft wool. It was established in Spain near the end of the Middle Ages, and was for several centuries kept as a strict Spanish monopoly; exports of the breed ...
is long and very soft. Coarse wools, being durable and resistant to pilling, are used for making tough garments and carpets.
Bone meal Bone meal is a mixture of finely and coarsely ground animal bones and slaughter-house waste products. It is used as a dietary supplement to supply calcium (Ca) and phosphorus (P) to monogastric livestock in the form of hydroxiapathite. As a slow-r ...
is an important fertilizer rich in calcium, phosphorus, and nitrogen, effective in removing
soil acidity Soil pH is a measure of the acidity or basicity (alkalinity) of a soil. Soil pH is a key characteristic that can be used to make informative analysis both qualitative and quantitatively regarding soil characteristics. pH is defined as the neg ...
. Bovid horns have been used as drinking vessels since antiquity.


In human culture

Bovidae have featured in stories since at least the time of
Aesop's fables Aesop's Fables, or the Aesopica, is a collection of fables credited to Aesop, a slave and storyteller believed to have lived in ancient Greece between 620 and 564 BCE. Of diverse origins, the stories associated with his name have descended to m ...
from
Ancient Greece Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cult ...
around 600 BC. Fables by Aesop include ''
The Crow and the Sheep The Crow and the Sheep is one of Aesop's Fables and is numbered 553 in the Perry Index. Only Latin versions of it remain. A sheep reproaches a crow that has perched on its back: 'If you had treated a dog in this way, you would have had your des ...
'', '' The Frog and the Ox'', and ''
The Wolf and the Lamb The Wolf and the Lamb is a well-known fable of Aesop and is numbered 155 in the Perry Index. There are several variant stories of tyrannical injustice in which a victim is falsely accused and killed despite a reasonable defence. The fable and i ...
''. The mythological creature Chimera, depicted as a lion, with the head of a goat arising from its back, and a tail that might end with a snake's head, was one of the offspring of Typhon and
Echidna Echidnas (), sometimes known as spiny anteaters, are quill-covered monotremes (egg-laying mammals) belonging to the family Tachyglossidae . The four extant species of echidnas and the platypus are the only living mammals that lay eggs and the ...
and a sibling of such monsters as
Cerberus In Greek mythology, Cerberus (; grc-gre, Κέρβερος ''Kérberos'' ), often referred to as the hound of Hades, is a multi-headed dog that guards the gates of the Underworld to prevent the dead from leaving. He was the offspring of the mo ...
and the
Lernaean Hydra The Lernaean Hydra or Hydra of Lerna ( grc-gre, Λερναῖα Ὕδρα, ''Lernaîa Hýdra''), more often known simply as the Hydra, is a serpentine water monster in Greek and Roman mythology. Its lair was the lake of Lerna in the Argolid, w ...
. The sheep, synonymous with the goat in Chinese mythology, is the eighth animal of the
Chinese zodiac The Chinese zodiac is a traditional classification scheme based on the lunar calendar that assigns an animal and its reputed attributes to each year in a repeating twelve-year cycle. Originating from China, the zodiac and its variations remai ...
, and a symbol of filial piety.


References


External links

* * {{Authority control Mammal families Extant Burdigalian first appearances Taxa named by John Edward Gray