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The black woodpecker (''Dryocopus martius'') is a large
woodpecker Woodpeckers are part of the bird family (biology), family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extrem ...
that lives in mature forests across the northern
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Si ...
. It is the sole representative of its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
in that region. Its range is expanding. The black woodpecker is easily the largest woodpecker species in Europe as well as in the portion of Asia where it lives and is one of the largest species worldwide. This non-migratory species tends to make its home in
old-growth forest An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological featur ...
or large forest stands and excavates a large tree hole to reside in. In turn, several species rely on black woodpeckers to secondarily reside in the holes made in trees by them. This woodpeckers diet consists mostly of
carpenter ant Carpenter ants (''Camponotus'' spp.) are large () ants indigenous to many forested parts of the world. They build nests inside wood consisting of galleries chewed out with their mandibles or jaws, preferably in dead, damp wood. However, unl ...
s. This species is closely related to, and fills the same
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...
in Europe as, the
pileated woodpecker The pileated woodpecker (''Dryocopus pileatus'') is a large, mostly black woodpecker native to North America. An insectivore, it inhabits deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the ...
of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the C ...
and the lineated woodpecker of South America, also being similar to the
white-bellied woodpecker The white-bellied woodpecker or great black woodpecker (''Dryocopus javensis'') is found in evergreen forests of tropical Asia, including the Indian subcontinent and Southeast Asia. It has 14 subspecies, part of a complex including the Andaman w ...
which is distributed to the south somewhat of the black woodpecker in Asia.


Taxonomy

The black woodpecker was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nom ...
'' under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Picus martius''. Linnaeus gave the locality as Europe, but this is now taken to be Sweden. The black woodpecker is now placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
'' Dryocopus'' that was introduced by the German naturalist
Friedrich Boie Friedrich Boie (4 June 1789 – 3 March 1870) was a German entomologist, herpetologist, ornithologist, and lawyer.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins Universit ...
in 1826. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''D. m. martius'' (Linnaeus, 1758) – western Europe to the
Kamchatka Peninsula The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
and Japan * ''D. m. khamensis'' ( Buturlin, 1908 – Tibet and southwestern China


Description

The black woodpecker measures long with a wingspan.''Woodpeckers: An Identification Guide to the Woodpeckers of the World'' by Hans Winkler, David A. Christie & David Nurney. Houghton Mifflin (1995), Body weight is approximately on average. Among standard measurements, the wing chord is , the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, ...
is , the very long bill is and the tarsus is . It is easily the largest woodpecker in its range and is second in size only to the great slaty woodpecker amongst the woodpecker species certain to exist (with the likely
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
of the
largest Large means of great size. Large may also refer to: Mathematics * Arbitrarily large, a phrase in mathematics * Large cardinal, a property of certain transfinite numbers * Large category, a category with a proper class of objects and morphisms (or ...
and second largest woodpeckers), although its average mass is similar to that of the Magellanic woodpecker of South America. The closely related pileated and
white-bellied woodpecker The white-bellied woodpecker or great black woodpecker (''Dryocopus javensis'') is found in evergreen forests of tropical Asia, including the Indian subcontinent and Southeast Asia. It has 14 subspecies, part of a complex including the Andaman w ...
s also broadly overlap in size with the black woodpecker, but both are somewhat smaller in average and maximal size and mass. The
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of this crow-sized woodpecker is entirely black apart from a red crown. In males, the entire crown is red, but in females only the top hindcrown is red, with the rest of the body all black. The juvenile black woodpecker is similar but is less glossy, with a duller red crown and a paler grey throat and bill .Perrins, C.M., Attenborough, D. and Arlott, N. (1987). ''New Generation Guide to the Birds of Britain and Europe''. University of Texas Press, Texas. The piercing yellow eyes and manic, high-pitched calls of the black woodpecker have made it the villain of fairy tales throughout its range. Their voice is remarkable in that it has two different calls. One is a short single high-pitched note, a loud, whistling ''kree-kree-kree'', done only twice in a row. The other is a screech-like shrill while in flight. Unlike other woodpecker species, the black woodpecker does not have a dipping, bounding flight, but instead flies with slow, unsteady-seeming wing beats with its head raised.


Distribution and habitat

The range of the black woodpecker spreads east from
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
across the whole of Europe, excluding
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
,
Ireland Ireland ( ; ga, Éire ; Ulster-Scots: ) is an island in the North Atlantic Ocean, in north-western Europe. It is separated from Great Britain to its east by the North Channel, the Irish Sea, and St George's Channel. Ireland is the s ...
, and northern
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and Swed ...
. It is also native to parts of
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
, including
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic of ...
,
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
and China, and to the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europea ...
, including
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkme ...
and
Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbekis ...
. The southern limits of this woodpecker's range are in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
and Italy, and it has also been recorded as a vagrant in
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a Sovereign state, country whose mainland is located on the Iberian Peninsula of Southern Europe, Southwestern Europe, and whose territory also includes ...
. The species is generally more uncommon and more discontinuous in distribution in the Asian part of its range.Peterson, R.T., Mountfort, G. and Hollom, P.A.D. (1993) ''Collins Field Guide: Birds of Britain and Europe''. HarperCollins Publishers, London. The black woodpecker is mainly found in forested regions, with a preference for extensive, mature woodland, including
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
,
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
, subtropical and
boreal forests Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, ...
. It is very widespread throughout mountainous and lowland forests. It is more likely to occur in marginal woods near human habitations during the non-breeding season. This species has been observed at elevations between . The black woodpecker is noticeably absent from the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
. Approximately 80 sightings of the species in the UK have been reported, but some of these are disputed, though the proximity of the British Isles to the species' range in Western Europe means that the species may cross over on a regular basis.


Behaviour and ecology

The woodpecker feeds by using its bill to hammer on dead trees to dig out
carpenter ant Carpenter ants (''Camponotus'' spp.) are large () ants indigenous to many forested parts of the world. They build nests inside wood consisting of galleries chewed out with their mandibles or jaws, preferably in dead, damp wood. However, unl ...
s and wood-boring beetle grubs. The selection of foods is relatively predictable, narrow and consistent in this species. Like all woodpeckers, this species has a specially adapted neck containing very strong muscles, which allow it to endlessly hack away at tree bark. Due to the size of its bill and large size and great physical power of this bird, it can access prey fairly deep within a tree. In order to position itself correctly, it has short, stumpy legs, as well as long, sharp claws and very stiff tail feathers. The woodpecker will more than likely choose for its nest a tree with a fungal disease, such as heart rot, although some will utilise a living, healthy tree. Once a hole has been made, the black woodpecker chips downwards through the trunk of the tree, creating a nesting chamber, the only lining being the woodchips created throughout the process. The black woodpecker's excavations provide homes for many other species of bird and mammal, and is therefore considered to be a "keystone" species in many of its habitats throughout its range. It not only provides habitats for other species, but also controls populations of wood-boring insects, helping to protect the trees. When the nest is ready, the female lays a single clutch of two to eight eggs, the average being four to six. The nest hole is usually dug in a live poplar or pine tree. The breeding pair take it in turns to incubate the eggs, also sharing duties of feeding and brooding the chicks once they have hatched. The nestlings may fight their way to the entrance of the nest in order to be fed first. After 18 to 35 days, the young black woodpeckers will leave the nest, staying with the adults for another week.


Status

The black woodpecker is a fairly widely distributed woodland species and can successfully breed in most areas where extensive woodland is left. At one point, when much of Europe and Asia was deforested, this species declined and in some areas is still struggling today, including in the Pyrenees. They normally require mature trees and ample stands of dead trees to sustain a viable breeding population. However, with the restoration of some forested areas, black woodpeckers have increased in some parts of Europe. They are occasionally considered a nuisance species due to their damage to power lines, communication poles and houses, occasionally resulting in woodpecker mortality due to electrocution or being culled by humans. The main cause of nesting failures appears to be predation. Their main natural predator is the
pine marten The European pine marten (''Martes martes''), also known as the pine marten, is a mustelid native to and widespread in most of Europe, Asia Minor, the Caucasus and parts of Iran, Iraq and Syria. It is listed as Least Concern on the IUCN Red List ...
(''Martes martes''), which feeds on eggs, nestlings and brooding females and then often takes over the nest hole of the woodpeckers for its own. Other than the marten, there are notably few known natural predators of black woodpeckers.
Western jackdaw The western jackdaw (''Coloeus monedula''), also known as the Eurasian jackdaw, the European jackdaw, or simply the jackdaw, is a passerine bird in the crow family. Found across Europe, western Asia and North Africa; it is mostly resident, al ...
s (''Corvus monedula'') are notably regular usurpers of this species' nest holes and a potential predator of eggs and small nestlings. A few of the larger birds of prey that can hunt in woodlands may prey on black woodpeckers. Among those recorded are Ural owls (''Strix uralensis''),
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and female ...
s (''Bubo bubo''),
northern goshawk The northern goshawk (; ''Accipiter gentilis'') is a species of medium-large raptor in the family Accipitridae, a family which also includes other extant diurnal raptors, such as eagles, buzzards and harriers. As a species in the genus '' Accip ...
s (''Accipiter gentilis''),
common buzzard The common buzzard (''Buteo buteo'') is a medium-to-large bird of prey which has a large range. A member of the genus '' Buteo'', it is a member of the family Accipitridae. The species lives in most of Europe and extends its breeding range acr ...
s (''Buteo buteo'') and
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds ...
s (''Aquila chrysaetos'').


Cultural significance

The municipality of
Nurmijärvi Nurmijärvi () is the most populated rural municipality of Finland, located north of the capital Helsinki. The neighboring municipalities of Nurmijärvi are Espoo, Vantaa, Tuusula, Hyvinkää and Vihti, and it is part of the Greater Helsinki ...
in
Uusimaa Uusimaa (; sv, Nyland, ; both lit. 'new land') is a region of Finland. It borders the regions of Southwest Finland, Tavastia Proper (Kanta-Häme), Päijänne Tavastia (Päijät-Häme), and Kymenlaakso. Finland's capital and largest city, He ...
,
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Both ...
has adopted the black woodpecker as the title bird of the municipality, because in addition to being the most common bird in the locality, it also appears in the literature of
Aleksis Kivi Aleksis Kivi (; born Alexis Stenvall; 10 October 1834 – 31 December 1872) was a Finnish author who wrote the first significant novel in the Finnish language, '' Seitsemän veljestä'' ("Seven Brothers") in 1870. He is also known for his 1864 ...
, a Finnish national author, originally from the Nurmijärvi. Nurmijärvi's local football club NJS has also adopted the black woodpecker as the club's logo. Dryocopus martius martius is thought to be the woodpecker referred to in the augural instructions on the early Italic
Iguvine Tablets The Iguvine Tablets, also known as the Eugubian Tablets or Eugubine Tables, are a series of seven bronze tablets from ancient Iguvium (modern Gubbio), Italy, written in the ancient Italic language Umbrian. The earliest tablets, written in the n ...
by the
Umbrian Umbrian is an extinct Italic language formerly spoken by the Umbri in the ancient Italian region of Umbria. Within the Italic languages it is closely related to the Oscan group and is therefore associated with it in the group of Osco-Umbrian ...
word ''peiqu'', a bird "very prominent in early Italic religion and mythology."Poultney, J.W. "Bronze Tables of Iguvium" 1959 p. 1 https://archive.org/details/bronzetablesofig00poul/page/n19/mode/2up


See also

* Black-bodied woodpecker *
Pileated woodpecker The pileated woodpecker (''Dryocopus pileatus'') is a large, mostly black woodpecker native to North America. An insectivore, it inhabits deciduous forests in eastern North America, the Great Lakes, the boreal forests of Canada, and parts of the ...


References


Bibliography

* Gorman, Gerard (2004): ''Woodpeckers of Europe: A Study of the European Picidae''. Bruce Coleman, UK. . * Gorman, Gerard (2011): ''The Black Woodpecker: A monograph on ''Dryocopus martius'' ''. Lynx Edicions, Barcelona. . {{Authority control black woodpecker Birds of Eurasia Birds of Russia black woodpecker black woodpecker