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''Batasio'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial no ...
of
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive ...
of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideal ...
Bagridae The Bagridae are a family of catfish that are native to Africa (''Bagrus'') and Asia (all other genera) from Japan to Borneo. It includes about 245 species. These fish are commonly known as naked catfishes or bagrid catfishes. Many large bagrids ...
. These small fish are found in fast-flowing hillstreams throughout
South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþaz' ...
and mainland
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
.


Taxonomy

''Batasio'' was first described by
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...
in 1860 with ''Batasio buchanani'' (a needless renaming of ''Batasio batasio'') designated as type. ''B. pakistanicus'', described in 1989, is tentatively placed in ''Batasio'', but its generic placement is questionable.


Species

There are currently 17 recognized species in this genus: * '' Batasio affinis'' Blyth, 1860 * '' Batasio batasio'' ( F. Hamilton, 1822) * '' Batasio convexirostrum'' Darshan, Anganthoibi & Vishwanath, 2011 * '' Batasio dayi'' ( Vinciguerra, 1890) * '' Batasio elongatus'' H. H. Ng, 2004 * '' Batasio fasciolatus'' H. H. Ng, 2006 * '' Batasio feruminatus'' H. H. Ng & Kottelat, 2008 * '' Batasio flavus'' Plamoottil, 2015 * '' Batasio fluviatilis'' ( F. Day, 1888) * '' Batasio macronotus'' H. H. Ng & Edds, 2004 * '' Batasio merianiensis'' ( B. L. Chaudhuri, 1913) * '' Batasio pakistanicus'' Mirza & M. A. Jan, 1989 * '' Batasio procerus'' H. H. Ng, 2008 * '' Batasio sharavatiensis'' Bhatt & Jayaram, 2004 * '' Batasio spilurus'' H. H. Ng, 2006 * '' Batasio tengana'' ( F. Hamilton, 1822) * '' Batasio tigrinus'' H. H. Ng & Kottelat, 2001


Distribution

''Batasio'' species are generally found throughout
South South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþaz' ...
and mainland
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical south-eastern region of Asia, consisting of the regions that are situated south of mainland ...
. ''B. affinis'' is found in
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
. ''B. fluviatilis'' is found in Northern
Malay Peninsula The Malay Peninsula ( Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The are ...
. ''B. tigrinus'' is also recorded from Thailand. ''B. dayi'' originates from the
Salween , ''Mae Nam Salawin'' ( , name_etymology = , image = Sweet_View_of_Salween_River_in_Tang_Yan_Township,_Shan_State,_Myanmar.jpg , image_size = , image_caption = Salween River in Shan State, Myanmar , map ...
and
Irrawaddy River The Irrawaddy River ( Ayeyarwady River; , , from Indic ''revatī'', meaning "abounding in riches") is a river that flows from north to south through Myanmar (Burma). It is the country's largest river and most important commercial waterway. Orig ...
drainages in Myanmar. ''B. elongatus'' is found in southwestern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
. ''B. macronotus'' originates from eastern
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
. ''B. pakistanicus'' is found in the
Indus River The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmir ...
. ''B. batasio'' also originates from northern
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
where it shares its habitat with '' Mystus vittatus''. ''B. fasciolatus'' is known from the Tista River drainage, a tributary of the
Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. ...
. ''B. merianiensis'' is found in the Brahmaputra River drainage. ''B. sharavatiensis'' originates from the
Sharavati River Sharavati is a river which originates and flows entirely within the state of Karnataka in India. It is one of the few westward flowing rivers of India and a major part of the river basin lies in the Western Ghats. The famous Jog Falls, located ...
in
Uttara Kannada Uttara Kannada is a district in the Indian states and territories of India, state of Karnataka. Uttara Kannada District is a major coastal district of Karnataka, and currently holding the title of the largest district in Karnataka. It is borde ...
,
Karnataka Karnataka (; ISO: , , also known as Karunāḍu) is a state in the southwestern region of India. It was formed on 1 November 1956, with the passage of the States Reorganisation Act. Originally known as Mysore State , it was renamed ''Karn ...
. ''B. spilurus'' is known from the Brahmaputra River drainage in the vicinity of
Dibrugarh Dibrugarh (pron: ˌdɪbru:ˈgɑ:) is an industrial city in Upper Assam with sprawling tea gardens. It is located 435 kms East from the state capital of Dispur. It serves as the headquarters of Dibrugarh district in the state of Assam ...
. ''B. tengana'' is known from the
Ganges The Ganges ( ) (in India: Ganga ( ); in Bangladesh: Padma ( )). "The Ganges Basin, known in India as the Ganga and in Bangladesh as the Padma, is an international river to which India, Bangladesh, Nepal and China are the riparian states." is ...
and the Brahmaputra River drainages; it is apparently restricted to upper reaches of larger rivers. ''B. travancoria'' is distributed in the Chittar, Kallada and
Pamba River The Pamba River (also called Pampa River) is the longest river in the Indian state of Kerala after Periyar and Bharathappuzha, and the longest river in the erstwhile former princely state of Travancore. Sabarimala temple dedicated to Lord Ayya ...
s in southern
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South C ...
and is considered a rare species.


Appearance and anatomy

''Batasio'' species typically have enlarged sensory pores on their head, laterally compressed heads and bodies, and at least 35
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e. ''B. tengana'', ''B. pakistanicus'' and ''B. spilurus'' differ from other ''Batasio'' species in that their adipose fin is much shorter. These fish are small catfish. ''B. pakistanicus'' and ''B. spilurus'' are the smallest species, reaching only SL and SL respectively. The other species range from SL. ''B. affinis'', ''B. dayi'', and ''B. fluviatilis'' have bars or bands, spots or stripes, and are not plain-bodied. ''B. pakistanicus'' has a black
humeral spot Humeral spot (from Latin ''humerus'', pertaining to the shoulder) is a mark or pattern found on several species of fish, typically above the pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding f ...
with a dark streak on the dorsum. ''B. batasio'' has a dark stripe along the
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
and another above. ''B. batasio'' does not exceed 10 centimetres (4 in). ''B. fasciolatus'' and ''B. tigrinus'' are the only ''Batasio'' species in which the adult colouration consists of five or six vertical dark brown bars on a light brown body. ''B. tengana'' has bands descending from the dorsal surface to the lateral line. ''B. travancoria'' has a lateral stripe along the lateral line and a faint shoulder spot. ''B. sharavatiensis'' is the only ''Batasio'' that has a completely plain and colourless body; its adipose fin is almost confluent (continuous) with the
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as s ...
save for a small notch. ''Batasio'', like most other bagrids, are easy to sex. The male has a visible
genital papilla The genital papilla is an anatomical feature of the external genitalia of some animals. In mammals In mammals, the genital papilla is a part of female external genitalia not present in humans, which appears as a small, fleshy flab of tissue. The p ...
just fore of the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
.
Gravid In biology and human medicine, gravidity and parity are the number of times a woman is or has been pregnant (gravidity) and carried the pregnancies to a viable gestational age (parity). These terms are usually coupled, sometimes with additional t ...
female ''B. tengana'' (South East Asian form) are easy to identify because the pink eggs can be seen through their semi-transparent belly when they swim near light.


Ecology

''Batasio'' species generally are found in fast-flowing hillstreams. ''B. fluviatilis'' is found in rivers and streams with moderate to swift current and a predominantly rocky bottom, and is less often in slow-flowing streams with a muddy substrate. ''B. tengana'' occurs in rivers and canals and is found in torrential streams. ''B. travancoria'' occurs in hillstreams and rivers at the bases of hills. ''B. fluviatilis'' hide among stones or submerged vegetation during the day and come out at night to feed. ''B. batasio'' appears to
mimic MIMIC, known in capitalized form only, is a former simulation computer language developed 1964 by H. E. Petersen, F. J. Sansom and L. M. Warshawsky of Systems Engineering Group within the Air Force Materiel Command at the Wright-Patterson AFB in ...
'' Mystus vittatus'', with which it inhabits the same habitat.


In the aquarium

''B. tengana'', though uncommon, is the most commonly encountered species of ''Batasio'' in the
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
hobby. ''B. tengana'' does well in captivity, and has been bred in captivity. The fish spawned in soft neutral water, but raising the fry, of which there were over 200, proved difficult. The temperature for the Indian species should remain between ; the Southeast Asian ''Batasio'' will tolerate slightly warmer waters. It appears that all ''Batasio'' need a lot of current and a high oxygen content. ''B. batasio'' is another species in the hobby, but can be easily confused with ''Mystus vittatus''. ''B. travancoria'' has only once been imported to the U.S. In captivity, the experience of one aquarist is that specimens of ''B. travancoria'' show little interest in all foods offered and slowly lose weight over a span of a few months and perish.


References

{{Taxonbar, from=Q3346558 Bagridae Fish of South Asia Fish of Southeast Asia Catfish genera Freshwater fish genera Taxa named by Edward Blyth