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''Butia eriospatha'' is a small species of ''
Butia ''Butia'' is a genus of palms in the family Arecaceae, native to the South American countries of Brazil, Paraguay, Uruguay and Argentina. Many species produce edible fruits, which are sometimes used to make alcoholic beverages and other foods. ...
''
palm Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the highlands of southern Brazil. It is very similar to ''B. odorata'', but is easily distinguished from this species by the distinct spathes which are densely covered in rust-coloured, woolly hairs. Indeed, the specific epithet is derived from Greek ''ἔριον'', wool, and Latin ''spatha'', which refers to the spathe. It has been given the name woolly jelly palm (UK) or wooly jelly palm (US) in English. Vernacular names for it where it is native are ''butiá-da-serra'', ''butiázeiro'', ''butiá-veludo'', ''butiá'' ''butiá verdadeiro'', ''butiá-do-campo'', ''yatáy'' and ''macumá''.


Taxonomy

In 1970 Sidney Fredrick Glassman moved this species, along with all other ''Butia'', to '' Syagrus'', but in 1979 he changed his mind and moved everything back.


Description

''Butia eriospatha'' is a solitary-trunked palm tree. The trunk is sometimes inclined to a side, and may occasionally be subterranean. The 20-22
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves arch back down towards the trunk and have a petiole armed with teeth spaced along their margins; the rachis of the leaf is 150–220 cm in length. The branched inflorescence develops in a woody, 115–135 cm long spathe covered in a dense woolly indumentum. The
pistillate Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
(female) flowers are 5–9mm in length. The shape of the fruit is globose (round), as is the shape of the nuts. Like all species of ''Butia'' studied, this species has relatively larger pollen grains than that of other genera of palm present in
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
, Brazil. These grains are bilaterally symmetrical, prolate-spheroid, monosulcate, and with the end piriform (pear-shaped). The surface is covered in minute 2μm-large reticulate patterns.


Distribution

It is native to southern Brazil, where the main bulk of its population is found straddling the highlands in the eastern Santa Catarina/
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
border regions, surrounded by scattered sub-populations in the southeastern coastal part of the high plateau of the states of Paraná, Rio Grande do Sul, and Santa Catarina. A naturalised population in an area of northern
Misiones Province Misiones (, ''Missions'') is one of the 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamia region. It is surrounded by Paraguay to the northwest, Brazil to the north, east and south, and Corrientes ...
, Argentina, was reported in 2008. Bauermann ''et al.'' investigated the possibility of using palm pollen, including this species, in
palynology Palynology is the "study of dust" (from grc-gre, παλύνω, palynō, "strew, sprinkle" and '' -logy'') or of "particles that are strewn". A classic palynologist analyses particulate samples collected from the air, from water, or from deposit ...
, in order to try to provide more detail about the ancient changes in habitat in the state Rio Grande do Sul in Brazil by tracking the changes in distribution and abundance of the palms, but were unable to provide much detail on the subject.


Habitat and ecology

It is found growing in the Atlantic rainforest
biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
on a plateau at higher altitudes, between 700 and 1,200m, where it is largely found in grasslands, aggregated in often extensive, dense, almost monoculture palm groves, but higher up also sometimes in open ''Araucaria'' forest. These distinctive dense palm groves are known as ''butiázais'' in
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
, but may also be known as ''butiatuba'' or ''butiazal''. The caterpillars of the butterflies ''Brassolis astyra'' ssp. ''astyra'' (''fide'' d'Aralljo & Silva ''et al.''
968 Year 968 ( CMLXVIII) was a leap year starting on Wednesday (link will display the full calendar) of the Julian calendar. Events By place Byzantine Empire * Emperor Nikephoros II receives a Bulgarian embassy led by Prince Boris (th ...
(=record no. 2362, ''Butia eriospatha'' identified as ''butiázeiro'' or ''Cocos'' sp. in this study)) and ''B. sophorae'' (possibly ssp. ''dinizi'') (=no. 2365) have been recorded feeding on ''B. eriospatha'' in Santa Catarina. ''
Opsiphanes invirae ''Opsiphanes'' is a genus of butterflies of the family Nymphalidae found from Mexico to South America. * ''Opsiphanes camena'' Staudinger, 886/small> * '' Opsiphanes cassiae'' (Linnaeus, 1758) * ''Opsiphanes cassina'' C. & R. Felder, 1862 – sp ...
'', the nominate form or possibly subspecies ''remoliatus'', was also recorded feeding on this palm in the same study (=no. 2390, 2392). The caterpillars of the Indonesian butterfly ''Cephrenes augiades'' ssp. ''augiades'' may also feed on the leaves this palm. In a long term study observing the feeding behaviour throughout the year of the squirrel ''Guerlinguetus brasiliensis'' ssp. ''ingrami'' in secondary ''
Araucaria angustifolia ''Araucaria angustifolia'', the Paraná pine, Brazilian pine or candelabra tree (, or ), is a critically endangered species in the conifer genus '' Araucaria''. Although the common names in various languages refer to the species as a "pine", it d ...
'' forest in the Parque Recreativo Primavera in the vicinity of the city of
Curitiba Curitiba () is the capital and largest city in the state of Paraná (state), Paraná in Brazil. The city's population was 1,948,626 , making it the List of cities in Brazil by population, eighth most populous city in Brazil and the largest in ...
, Paraná, of the ten plant species of which the squirrel ate the seeds or nuts, ''Butia eriospatha'' was an uncommon but regular part of the diet in the spring and summer. Mushrooms, seeds of ''Araucaria angustifolia'', nuts of '' Syagrus romanzoffiana'' (queen palm) and acorns of ''
Quercus robur ''Quercus robur'', commonly known as common oak, pedunculate oak, European oak or English oak, is a species of flowering plant in the beech and oak family, Fagaceae. It is a large tree, native plant, native to most of Europe west of the Caucasus ...
'' (English oak) were the main part of the diet depending on the season.


Uses

''Butia eriospatha'' is reasonably available in Europe as an ornamental. It is often used in mild temperate climates as a hardy palm for its exotic look. Products obtained from the palm are used locally and are not widely marketed. The fruit pulp is used as a base to make beverages. A wine is made from the palm, as are juices and jellies/jams. Fibres were once harvested from this palm, which were used to make mattresses. Plantations of this palm were planted in the 1950s in Brazil for this purpose.


Horticulture

It is sometimes cultivated in Brazil and Argentina. It is advised to plant the palms in full sunlight. It is said to take -14 °C, but should be protected at -6 °C in
the Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
.


Conservation

The species is widely cultivated, but according to Noblick in 1998, wild populations are declining. It was listed as ''em perigo'' (endangered) by the Rio Grande do Sul state government in 2002 (''decreto'' nº 42.099) and 2014 (''decreto'' n° 52.109), because the population is believed to have been reduced by over 50% in at least the last 100 years. In 2012 the
Centro Nacional de Conservação da Flora The Centro Nacional de Conservação da Flora (CNCFlora) is a Brazilian nonprofit organization that determines conservation statuses of various Brazilian plant species. It intends to create a Red List (''lista vermelha'' in Portuguese) of plants ...
rated the conservation status for Brazil as 'vulnerable'.


Gallery

File:Butia eriospatha.jpg, Note the teeth along the margins of the leaf petiole on this plant for sale in the USA. File:Butia eriospatha kz1.JPG, Young ''Butia eriospatha'' palm cultivated in the
Jardín Botánico Canario Viera y Clavijo Jardín Botánico Canario Viera y Clavijo is the full name of the botanical garden on Gran Canaria, one of the Canary Islands. "Jardín Botánico Canario" means "Botanical Garden of the Canaries", while the additional words "Viera y Clavijo" honor ...
, flowering at the end of March. File:Butia eriospatha kz2.JPG, Flowers of ''B. eriospatha'' at the end of March at the Jardín Botánico Canario Viera y Clavijo. File:Butiá.jpg, Ripe fruit in December (summer) in Brazil. Butia eriospatha - Jardim Botânico da Universidade de Coimbra - Coimbra, Portugal - DSC08890.jpg, Fruit on ground after ripening,
Botanical Garden of the University of Coimbra The Botanical Garden of the University of Coimbra ( or simply ''Jardim Botânico'') is a botanical garden in Coimbra, Portugal. History It was founded in 1772 by Sebastião José de Carvalho e Melo (the Marquis of Pombal). The location for ...
. File:Contributions du Jardin botanique de Rio de Janeiro (1901) (20692838501).jpg, Comparison of fruit by
João Barbosa Rodrigues João Barbosa Rodrigues (June 22, 1842 – March 6, 1909) was considered one of Brazil's greatest botanists, known especially for his work on orchids and palms. For nearly two decades he was director of the Botanic Garden of Rio de Janeiro. Somet ...
in 1901. ''B. eriospatha'' is 'D' (note the smaller, very round fruit); ''B. yatay'' is 'A', ''B. odorata'' is 'B' & 'C', and '' Syagrus coronata'' is 'E'.


References

{{Taxonbar, from=Q1935582 eriospatha Endemic flora of Brazil Flora of Paraná (state) Flora of Rio Grande do Sul Flora of Santa Catarina (state) Vulnerable flora of South America Taxa named by Odoardo Beccari Taxonomy articles created by Polbot