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''Bugtilemur'' is an extinct genus of Strepsirhine
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
belonging to the adapiform family Ekgmowechashalidae.It is represented by only one species, ''B. mathesoni'', which was found in the
Chitarwata Formation The Chitarwata Formation is a geological formation in western Pakistan, made up of Oligocene and early Miocene terrestrial fluvial facies. The sediments were deposited in coastal depositional environments (estuarine, strandplain and tidal flats) ...
of
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
. When first described, ''Bugtilemur'' was classified in the lemur family
Cheirogaleidae The Cheirogaleidae are the family of strepsirrhine primates containing the various dwarf and mouse lemurs. Like all other lemurs, cheirogaleids live exclusively on the island of Madagascar. Characteristics Cheirogaleids are smaller than the othe ...
, complicating the picture of the early evolution of lemurs by suggesting that lemurs originated in Asia. Described from a few teeth, the specimen possesses a lower canine that, according to Marivaux et al., confirms the presence of the strepsirrhine-specific
toothcomb A toothcomb (also tooth comb or dental comb) is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates ( ...
. Furthermore, on the basis of cheektooth morphology, the molars share strong affinities with those of the genus ''Cheirogaleus'' (
dwarf lemur The dwarf lemurs are the lemurs of the genus ''Cheirogaleus''. All of the species in this genus, like all other lemurs, are native to Madagascar. Description Measuring 19–27 cm in body length with a tail about 16–17 cm, they are ...
s). However, ''Bugtilemur'' appears to be much smaller than the
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
Malagasy genus and its toothcomb was shorter and broader. More recently, the structure and general presence of the toothcomb in ''Bugtilemur'' has been questioned, as well as many other dental features, suggesting that it is most likely an adapiform. The adapiform nature of ''Bugtilemur'' was confirmed in a 2016 cladistic analysis that recovered it in the family Ekgmowechashalidae, which is more consistent with the lemur fossil record.


See also

*
Karanisia ''Karanisia'' is an extinct genus of strepsirrhine primate from middle Eocene deposits in Egypt. Classification Two species are known, ''K. clarki'' and ''K. arenula''. Originally considered a crown lorisid, more comprehensive phylogenetic a ...


References

{{Taxonbar, from=Q926781 Prehistoric strepsirrhines Eocene primates Prehistoric primate genera Fossil taxa described in 2001 Eocene mammals of Asia