Buddleja Crispa
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''Buddleja crispa'', the Himalayan butterfly bush, is a deciduous shrub native to
Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bordere ...
,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
,
North India North India is a loosely defined region consisting of the northern part of India. The dominant geographical features of North India are the Indo-Gangetic Plain and the Himalayas, which demarcate the region from the Tibetan Plateau and Central ...
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Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
,
Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...
and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
(
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...
,
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
,
Tibetan Autonomous Region The Tibet Autonomous Region or Xizang Autonomous Region, often shortened to Tibet or Xizang, is a province-level autonomous region of the People's Republic of China in Southwest China. It was overlayed on the traditional Tibetan regions of Ü ...
), where it grows on dry river beds, slopes with boulders, exposed cliffs, and in thickets, at elevations of 1400–4300 m.Li, P. T. & Leeuwenberg, A. J. M. (1996). Loganiaceae, in Wu, Z. & Raven, P. (eds) ''Flora of China'', Vol. 15. Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis, USA.
online at www.efloras.org
/ref> Named by Bentham in 1835, ''B. crispa'' was introduced to cultivation in 1850,Stuart, D. (2006). ''Buddlejas''. RHS Plant Collector Guide. Timber Press, Oregon, US. and came to be considered one of the more attractive species within the genus; it ranked 8th out of 57 species and cultivars in a public poll organized by the Center for Applied Nursery Research (CANR) at the
University of Georgia , mottoeng = "To teach, to serve, and to inquire into the nature of things.""To serve" was later added to the motto without changing the seal; the Latin motto directly translates as "To teach and to inquire into the nature of things." , establ ...
, US.Gillman, J., Dirr, M. A. & Braman, K. (1997). ''Evaluation and selection of superior Buddleja taxa for Georgia nurseries and gardens.'' Center for Applied Nursery Research, Dearing, Georgia, USA. In the UK, ''B. crispa'' was accorded the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Merit The Award of Merit, or AM, is a mark of quality awarded to plants by the British Royal Horticultural Society (RHS). The award was instituted in 1888, and given on the recommendation of Plant Committees to plants deemed "of great merit for exhibitio ...
in 1961.Hillier & Sons. (1990). ''Hillier's Manual of Trees & Shrubs, 5th ed.''. p. 47. David & Charles, Newton Abbot. However, the species is not entirely cold-hardy, and thus its popularity is not as ubiquitous as it might otherwise be.


''Buddleja crispa'', after Leeuwenberg

In his 1979 revision of the taxonomy of the African and Asiatic species of Buddleja, the Dutch botanist Toon Leeuwenberg sank five Chinese species as ''B. crispa'' on the basis of the similarity in the individual flowers, dismissing the wide ranges in size of both
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
and
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
as attributable to environmental factors.Leeuwenberg, A. J. M. (1979) ''The Loganiaceae of Africa XVIII Buddleja L. II, Revision of the African & Asiatic species''. H. Veenman & Zonen B. V., Wageningen, Nederland. It was Leeuwenberg's taxonomy which was adopted in the ''Flora of China'' published in 1996. The five former species, still widely recognized in horticulture, are: '' Buddleja agathosma'', '' Buddleja caryopteridifolia'', '' Buddleja farreri'', '' Buddleja sterniana'', and ''
Buddleja tibetica ''Buddleja tibetica'' was a species Synonym_(taxonomy)#Botany, sunk as ''Buddleja crispa'' by Anthonius Josephus Maria Leeuwenberg, Leeuwenberg in 1979,Leeuwenberg, A. J. M. (1979) ''The Loganiaceae of Africa XVIII Buddleja L. II, Revision of the ...
''.


Description

The 'original' ''B. crispa'' as known to horticulture, cloned from a plant grown at
Aldenham Aldenham is a village and civil parish in Hertfordshire, north-east of Watford and southwest of Radlett. It was mentioned in the Domesday Book and is one of Hertsmere's 14 conservation areas. The village has eight pre-19th-century listed build ...
, England (see Cultivation), is a comparatively slow growing
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
of bushy habit, reaching 3.5 m high, more in spread.Bean 1976, vol 1, p. 450. ''The Flora of China'' gives a height of up to 5 m. Young twigs and both sides of the leaves are covered with a white or tawny loose felt. The leaves are ovate-lanceolate, 5–12 cm long by 2.0–4.5 cm wide, with petioles 0,6–2,5 cm. The shrub flowers from February to August.Flora of China; Bean says June or later; Krüssmann 1984, vol I, p. 242 says July - August; Hillier 1990, p. 47 says late summer; Phillips 1989, p. 209 says: flowers in spring - often before the leaves expand - if not pruned; summer to autumn if pruned in spring." The fragrant flowers form terminal
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s 7–10 cm long by 5 cm wide. The corolla is
lilac ''Syringa'' is a genus of 12 currently recognized species of flowering plant, flowering woody plants in the olive family or Oleaceae called lilacs. These lilacs are native to woodland and scrub from southeastern Europe to eastern Asia, and wid ...
, with an orange throat.A. Mehra et al., Occurrence of Chilli veinal mottle virus in Himalayan butterfly bush (Buddleja crispa)
accessed 21 November 2007
Ploidy Ploidy () is the number of complete sets of chromosomes in a cell (biology), cell, and hence the number of possible alleles for Autosome, autosomal and Pseudoautosomal region, pseudoautosomal genes. Sets of chromosomes refer to the number of mat ...
2''n'' = 38 (diploid).Chen, G, Sun, W-B, & Sun, H. (2007). Ploidy variation in Buddleja L. (Buddlejaceae) in the Sino - Himalayan region and its biogeographical implications. ''Botanical Journal of the Linnean Society.'' 2007, 154, 305 – 312. The Linnean Society of London. The former species sunk by Leeuwenberg, as listed in the preceding section, have, with the exception of "''B. sterniana''", inflorescences of varying density < 12  cm long, complemented by leaves of variable size and shape, often covered in a dense white tomentum when young. The exception, "''B. sterniana''", has markedly smaller inflorescences and leaves < 6  cm long.


Cultivation

''Buddleja crispa'' needs a well-drained soil and full sun;
Bean A bean is the seed of several plants in the family Fabaceae, which are used as vegetables for human or animal food. They can be cooked in many different ways, including boiling, frying, and baking, and are used in many traditional dishes th ...
states that it is at its best when grown on a wall.Bean shows a b/w-photo (Plate 23) of a beautiful plant, indeed growing against a wall Most if not all the specimens in commerce in the UK derive from a plant in the Aldenham collection amassed by
Vicary Gibbs Vicary Gibbs may refer to: * Vicary Gibbs (judge) (1751–1820), English barrister, judge and politician * Vicary Gibbs, 6th Baron Aldenham (born 1948), British peer * Vicary Gibbs (St Albans MP) Vicary Gibbs (12 May 1853 – 13 January 1932) w ...
.Bean, W. J. (1914). ''Trees and shrubs hardy in the British Isles''. Eighth edition, revised by D. L. Clarke, 1989. Vol. 1, A-C. Murray, London. Hardiness:
USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...
zones 8–9.


Notable specimens

A particularly tall example of tree-like form > 4.5 m high is grown near an entrance to the grassland and aquatic gardens at the
Royal Botanic Gardens, Kew Royal Botanic Gardens, Kew is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. An internationally important botanical research and education institution, it employs 1,100 ...
.


Cultivars

* 'Huimoon' = selling name images: a
Modeste Herwig's Photo Website
and a

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References


Literature

*Bean, W. J. (1970). ''Trees & Shrubs Hardy in the British Isles, 8th ed., Vol. 1.''. (2nd impression 1976) London *Hillier & Sons. (1990). ''Hillier's Manual of Trees & Shrubs, 5th ed.''. David & Charles, Newton Abbot. *Krüssmann, G. (1984). ''Manual of Cultivated Broad-leaved Trees & Shrubs, Vol. 1''. Engl. transl. London, 1984. *Phillips, R. & Rix, M. (1989). ''Shrubs'', Pan Books, London. *Stuart, D. (2006). ''Buddlejas''. Timber Press, Oregon, USA. {{Taxonbar, from=Q4984447 crispa Flora of Afghanistan Flora of China Flora of the Indian subcontinent Plants described in 1835