Brown Awl
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''Badamia exclamationis'', commonly known as the brown awl or narrow-winged awl,Markku Savela's website on Lepidoptera. Page on genu

TOL web page on genu
''Badamia''
/ref> is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
belonging to the family
Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
. It is found in
south South is one of the cardinal directions or Points of the compass, compass points. The direction is the opposite of north and is perpendicular to both east and west. Etymology The word ''south'' comes from Old English ''sūþ'', from earlier Pro ...
and
southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, and
Oceania Oceania (, , ) is a region, geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern Hemisphere, Eastern and Western Hemisphere, Western hemispheres, Oceania is estimated to have a land area of ...
.


Range

The brown awl is found in
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
, South
Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. This butterfly is found throughout the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
and in the
Andaman islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
. The type locality is
South India South India, also known as Dakshina Bharata or Peninsular India, consists of the peninsular southern part of India. It encompasses the Indian states of Andhra Pradesh, Karnataka, Kerala, Tamil Nadu, and Telangana, as well as the union territo ...
.


Status

As per William Harry Evans (1932), the butterfly is common in India and rare in the
Andaman islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
. Mark Alexander Wynter-Blyth (1957) records it as "Not Rare" and "Locally Common". Krushnamegh Kunte (2000) reports it as common in deciduous forests during the monsoon months and the evergreen forests in the following months.


Habits

A forest butterfly, the brown awl favours openings and edges of deciduous and evergreen forests while its caterpillars are to be found in moist deciduous and semi-evergreen forests. It flies about either late or early in the morning in the shade of the jungles. It can be sometimes seen in bright sunlight visiting flowers, such as ''
Glycosmis ''Glycosmis'' is a genus of flowering plants in the citrus family, Rutaceae and tribe Clauseneae.''Glyc ...
'', '' Buddleia'', ''
Chromolaena ''Chromolaena'' is a genus of about 165 species of perennials and shrubs in the family Asteraceae. The name is derived from the Greek language, Greek words (), meaning "color", and () or () meaning "cloak". It refers to the colored phyllaries ...
'' and '' Lantana'', but is very wary and energetic at such times, moving jerkily and rapidly between flowers or across
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s. It can also be seen mud-puddling or at bird droppings. During a population explosion, like those of the common banded awl ('' Hasora chromus''), the caterpillars of the brown awl may strip away all their food supply forcing the butterflies to migrate to other places where a fresh supply of host plants is available and even to other habitats such as shrubs, grasslands and gardens. The brown awl flies as low as 6 feet over the bushes or as high as 60 to 75 feet in the canopy. The adults feed at lower levels on flowers of shrubs and small trees, but ascend to higher reaches of the vegetation to lay eggs or to bask, which it does very occasionally, holding its wings flat with the forewings covering the hindwings thus giving an arrowhead effect. The flight of the butterfly is fast and bounding with an audible wing beat. When inactive, it rests on the undersides of leaves in shady forest spots, with the head pointing downwards. If disturbed it will generally buzz around energetically before returning to the same spot to rest.


Description

The brown awl is a non-descript brown butterfly, darker above and lighter below. The sexes are alike, except for three to four semi-transparent spots on the forewing which cannot be differentiated in the field. The skipper has a light-brown abdomen with black bands across it. The dry-season form is usually smaller, paler, and may not have the forewing spots. This skipper is unmistakable because of its long and narrow wings. It has the longest wings in proportion to breadth of all Indian butterflies.


Detailed description

Edward Yerbury Watson Edward Yerbury Watson (27 July 1864 – 8 November 1897) was an English entomologist who specialised in Lepidoptera most notably Hesperiidae. Yerbury Watson was born in British India to Joanna and George Edward Watson. He joined the North Lancash ...
(1891) gives a detailed description: Genus characters Species description


Similar species

Unlike the other awls, the brown awl lacks the narrow white wing bands on the hindwings. The very distinctive characteristics of the brown awl are the characteristic shape of the body and the narrower wings than the other awls.


Life cycle


Eggs

The brown awl lays many eggs on a single plant, one at a time, on the tips of fresh shoots. The dome-shaped egg is pale green with longitudinal ridges having fine beadings; a total 13 ridges in all.


Larva

The larva is a pale violaceous (violet) yellow, with numerous black transverse dorsal lines; the prolegs are whitish encircled with black. The head is yellow, approximately heart shaped, with a black band and many tiny black spots. On hatching the larva webs the edges of leaves together with silk to form a roomy cell from a leaf in which it resides throughout the larval stage. When disturbed, it can move quite briskly and even drop off. The caterpillars of the brown awl grow faster than most of those of other families, and have moist, sticky droppings. At the time of pupation they descend close to the ground, looking for suitable spots to pupate. The caterpillar constructs a tubular cell from a leaf by drawing the edges together with thick strands of silk. In this cell, the caterpillar prepares an extensive silk bed on which it sits awaiting pupation. The freshly formed pupa clings onto the silken pad almost immediately.


Pupa

The pupa is stubby, with protruding eyes and a prominent projection on the head in between them. The pupa may be light brown or violaceous. The body tapers away from the shoulders towards the rear. The abdomen is creamish with a row of four black spots on each side. The pupa is shiny, but plastered with a white powder.


Host plants

The caterpillars have been recorded on the following deciduous and semi-evergreen forest plants, mostly from the family Combretaceae: * Large climbing shrubs of '' Combretum'' such as ''
Combretum albidum ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...
'', ''
Combretum latifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...
'' and ''
Combretum ovalifolium ''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
''. * Large forest trees of genus ''
Terminalia Terminalia may refer to: * Terminalia (festival), a Roman festival to the god of boundaries Terminus * ''Terminalia'' (plant), a tree genus * Terminalia (insect anatomy), the terminal region of the abdomen in insects * ''Polyscias terminalia'', a ...
'' such as ''
Terminalia bellerica ''Terminalia bellirica'', known as baheda, bahera, behada, beleric or bastard myrobalan (Arabic: beliledj بليلج, borrowed from Middle Persian Balilag), Persian بلیله (Balileh), Sanskrit: Bibhitaka बिभीतक, Aksha is a large ...
'', '' Terminalia catappa'' and '' Terminalia oblongata''. * ''
Anogeissus acuminata ''Terminalia phillyreifolia'' is an Asian species of tree in the family Combretaceae. It has been called buttontree or yon (from ; IPA: ). It is a medium-sized tree found in both primary and secondary tropical and sub-tropical forests. It is reco ...
'' and '' Chionanthus purpureus''. * '' Hiptage benghalensis'' (
Malpighiaceae Malpighiaceae is a family of flowering plants in the order Malpighiales. It comprises about 73 genera and 1315 species, all of which are native to the tropics and subtropics. About 80% of the genera and 90% of the species occur in the New World ( ...
)nhm.ac.uk Caterpillar Host plant database
/ref> * '' Linociera purpurea''. * ''
Ficus ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending in ...
'' spp.


Gallery

image:Badamia exclamationis (ento-csiro-au).jpg, Brown awl Image:ExclamationisCRW 5480-01.jpg, Museum specimens File:7淡綠弄蝶5(劉威良攝) (31621207561).jpg, Dorsal view File:7淡綠弄蝶21(李榮芳攝) (30927073453).jpg, Mating pair


See also

*
Coeliadinae Coeliadinae is a subfamily of the skipper butterfly family (Hesperiidae). With about 150 described species, this is one of several smallish skipper butterfly subfamilies. It was first proposed by William Frederick Evans in 1937.Brower & Warren ( ...
*
List of butterflies of India (Coeliadinae) A total of 22 species belonging to four genera of the subfamily Coeliadinae (family Hesperiidae), or the awls, awlets and awlkings, as they are commonly called, are found in India. These are relatively large skippers which inhabit dense forests, ...
*
List of butterflies of India (Hesperiidae) India has a rich biodiversity of butterflies, of which skippers are a well represented family. Of the seven subfamilies belonging to the family Hesperiidae, four are found in India, comprising a total of 223 species of 74 genera and these are l ...


References


Bibliography

* * Brower, Andrew V. Z. and Warren, Andrew, (2007). Coeliadinae Evans 1937. Version 21 February 2007 (temporary). http://tolweb.org/Coeliadinae/12150/2007.02.21 in The Tree of Life Web Project, http://tolweb.org/ * * * * Watson, E. Y. (1891) ''Hesperiidae indicae''. Vest and Co. Madras. *


External links


Images representing ''Badamia exclamationis''
at Consortium for the Barcode of Life {{DEFAULTSORT:Badamia Exclamationis Coeliadinae Butterflies of Asia Butterflies of Oceania Butterflies of Indochina Butterflies of Malaysia Butterflies of Singapore Butterflies described in 1775 Taxa named by Johan Christian Fabricius