Brown Argus
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''Aricia agestis'', the brown argus, is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Lycaenidae Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...
. It is found throughout the
Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
, north to northern
Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
(
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...
) and east to Siberia and the
Tian Shan The Tian Shan,, , otk, 𐰴𐰣 𐱅𐰭𐰼𐰃, , tr, Tanrı Dağı, mn, Тэнгэр уул, , ug, تەڭرىتاغ, , , kk, Тәңіртауы / Алатау, , , ky, Теңир-Тоо / Ала-Тоо, , , uz, Tyan-Shan / Tangritog‘ ...
.


Subspecies

*''A. a. agestis'' southern and central Europe *''A. a. calida'' Chavignerie Sicily, Italy, Asia Minor *''A. a. azerbaidzhana'' Obraztsov, 1935 Transcaucasia, Caucaus Major *''A. a. nazira'' (Moore, 1865) Darvaz, western Pamirs, north-western Himalayas Brown argus (Aricia agestis) female.jpg, female ''A. a. agestis, UK Brown argus (Aricia agestis calida) Italy.jpg, female ''A. a. calida'', Italy Aricia agestis, Brown Argus, Minera, North Wales, May 2017 (34680423115).jpg , larva


Description in Seitz

''L. astrarche'' Bgstr. (= ''medon'' Hufn., ''agestis'' Schiff., ''idas'' Gerh. (?), ''nazira'' Moore) (79 k). Above similar to the preceding, L._anteros_.html" ;"title="Aricia_anteros.html" ;"title="Aricia anteros">L. anteros ">Aricia_anteros.html" ;"title="Aricia anteros">L. anteros deep dark brown, with strongly marked discocellular spot, especially in the male. Typical specimens have this spot black and bear red spots at the outer margin. Underside with numerous ocelli, the hindwing with a pale smear from the apex of the cell to the centre of the outer margin. From Scandinavia to the Sahara and northern India, and from the Canary Isles to the island of Askold in the Pacific, ascending in the mountains up to 10,000 ft. (Doherty). — Besides accidental aberrations in the usual directions of variation, a number of seasonal and geographical forms have been established. If the spring-brood of the Central European form is regarded as name-typical ''astrarche'', ab. ''aestiva'' Stgr. is the name for the second brood, which occurs regularly in the south and more rarely in the north and which connects ''astrarche'' with ''calida'' Bell. (79 k, 80 a) [''A. a. calida'' (Bellier, 1862) . The latter has a much darker, almost coffee-brown, under-side, with a broader band of red spots. Egg pale green, flattened, with the top concave and the surface minutely reticulate (Tutt, Harrison, Gillmer). Larva light green, with a purple dorsal stripe and a similar stripe along the sides, between them oblique pale smears; in spring and summer (in the south throughout the year) on various plants, such as ''Helianthemum'', ''Centaurea'', ''Erodium'', etc. Pupa pale yellowish green, sometimes with a brown dorsal stripe, on or near the ground. The butterflies are on the wing in May and again from July onward, in the south the whole summer until the autumn, in several broods. They fly everywhere, in fields, on field-paths, on rocky hills, even in gardens, always close to the ground and belong to the very commonest butterflies. During the hours of flight they settle with spread wings on blades of grass, blossoms, or clods of earth, while they sleep with closed wings generally hidden under umbels and the inflorescences of grasses.


Appearance and biology (Great Britain)

:''Note that information on this species applies to Great Britain and some details may not be consistent with the species in other parts of its range.'' Although one of the "blues", both sexes are brown on the uppersides with a band of orange spots at the border of each wing. They can be mistaken for other female blues but the brown argus never has any blue scales at the base of the wings like other female blues often do. It could also be mistaken for the northern brown argus (''Aricia artaxerxes'') were it not for the fact that their ranges do not overlap in the UK, unlike on continental Europe. The underside has the typical "blue" pattern of a greyish/brownish ground colour with black spots outlined in white and a row of orange spots along the border. The pattern of the black spots is the best way to distinguish this species from female
common Common may refer to: Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Common, common land area in Cambridge, Massachusetts * Clapham Common, originally com ...
, chalkhill and
Adonis In Greek mythology, Adonis, ; derived from the Canaanite word ''ʼadōn'', meaning "lord". R. S. P. Beekes, ''Etymological Dictionary of Greek'', Brill, 2009, p. 23. was the mortal lover of the goddess Aphrodite. One day, Adonis was gored by ...
blues as they lack the black spot found near the base of the forewing which is present on these three species. This species has seen an expansion in its range in recent years and is widely distributed across south-east England and most of
the Midlands The Midlands (also referred to as Central England) are a part of England that broadly correspond to the Kingdom of Mercia of the Early Middle Ages, bordered by Wales, Northern England and Southern England. The Midlands were important in the Ind ...
with colonies occurring in
Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
and as far north as
Yorkshire Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a Historic counties of England, historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other Eng ...
. These northern sites have seen a lot of confusion in recent years with genetic studies looking at various colonies to separate the two ''Aricia'' species. Until a few years ago, these northern colonies were thought to be the northern brown argus and more colonies may yet be found to be misidentified. Like other blues it is common on the
chalk downland Downland, chalkland, chalk downs or just downs are areas of open chalk hills, such as the North Downs. This term is used to describe the characteristic landscape in southern England where chalk is exposed at the surface. The name "downs" is deriv ...
s of southern England but will also use other habitats such as woodland clearings, coastal grasslands and heathland. It is not closely related to the
Scotch argus The Scotch argus (''Erebia aethiops'') is a butterfly of the family Nymphalidae. In spite of its English name ''argus'', it is not a close relation of the brown argus nor the Aricia artaxerxes, northern brown argus. Taxonomy Subspecies include: ...
.


Life cycle and food plants

:''Note that information on this species applies to Great Britain and some details may not be consistent with the species in other parts of its range.'' Common rock-rose (''
Helianthemum nummularium ''Helianthemum nummularium'' (known as common rock-rose) is a species of rock-rose (Cistaceae), native to most of Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical si ...
'') is the favoured food plant on
calcareous soil Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
s. In other habitats dove's-foot cranesbill ('' Geranium molle'') and common stork's-bill (''
Erodium cicutarium ''Erodium cicutarium'', also known as common stork's-bill, redstem filaree, redstem stork's bill or pinweed, is a herbaceous annual plant, annual – or in warm climates, biennial plant, biennial – member of the family Geraniaceae of flowering ...
'') are used and possibly other '' Geranium'' species as well. Eggs are laid singly on the underside of leaves. The typically
slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a smal ...
-like lycid larvae are green with a pale line along each side and always attended by ants. They hibernate as fully-grown larvae and pupate the following spring. There are two broods a year in the southern colonies with adults on the wing in May and June and again in late July till mid-September but further north they are single brooded and fly in June and July. Learn about butterflies
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See also

*
List of butterflies of Great Britain This is a list of butterflies of Great Britain, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in ''The Moths and Butterflies of Great Britain and Ireland'' by Emmet ''et al.'' and '' ...


References

{{Taxonbar, from=Q1294181 agestis Butterflies of Europe