Bronze Mannikin
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The bronze mannikin or bronze munia (''Spermestes cucullata'') is a small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
(i.e. perching)
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
of the
Afrotropics The Afrotropical realm is one of Earth's eight biogeographic realms. It includes Africa south of the Sahara Desert, the majority of the Arabian Peninsula, the island of Madagascar, southern Iran and extreme southwestern Pakistan, and the island ...
. This very social estrildid finch is an uncommon to locally abundant bird in much of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
south of the Sahara Desert, where it is resident, nomadic or irruptive in mesic savanna or
forest margin A woodland edge or forest edge is the transition zone (ecotone) from an area of woodland or forest to fields or other open spaces. Certain species of plants and animals are adapted to the forest edge, and these species are often more familiar to hu ...
habitats. It has an estimated global extent of occurrence of 8,100,000 km2. It is the smallest and most widespread of four munia species on the African mainland, the other being
black-and-white Black-and-white (B&W or B/W) images combine black and white in a continuous spectrum, producing a range of shades of grey. Media The history of various visual media began with black and white, and as technology improved, altered to color. ...
, red-backed and magpie mannikin. It co-occurs with the Madagascar mannikin on the
Comoro Islands The Comoro Islands or Comoros ( Shikomori ''Komori''; ar, جزر القمر , ''Juzur al-qamar''; french: Les Comores) form an archipelago of volcanic islands situated off the southeastern coast of Africa, to the east of Mozambique and northw ...
, and was introduced to
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...
. Especially in the
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
, it is considered a pest in grain and rice fields. It is locally trapped for the pet bird trade.


Taxonomy

The bronze mannikin was formally described in 1837 by the English naturalist
William John Swainson William John Swainson FLS, FRS (8 October 1789 – 6 December 1855), was an English ornithologist, malacologist, conchologist, entomologist and artist. Life Swainson was born in Dover Place, St Mary Newington, London, the eldest son of ...
and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Spermestes cucullata''. The specific epithet is from
Late Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in t ...
''cucullatus'' meaning "hooded". The type locality is Senegal. Two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''S. c. cucullata'' Swainson, 1837 – Senegal and Gambia to south Sudan, Uganda, west Kenya and the island of Bioko * ''S. c. scutata''
Heuglin Martin Theodor von Heuglin (20 March 1824, Hirschlanden, Württemberg5 November 1876), was a German explorer and ornithologist. Biography Heuglin was born in Hirschlanden (now part of Ditzingen) in Württemberg. His father was a Protestant past ...
, 1863 – Ethiopia and east Kenya to southeast South Africa, east Angola and
Comoro Islands The Comoro Islands or Comoros ( Shikomori ''Komori''; ar, جزر القمر , ''Juzur al-qamar''; french: Les Comores) form an archipelago of volcanic islands situated off the southeastern coast of Africa, to the east of Mozambique and northw ...


Description

The bronze mannikin is one of the smallest munia species,
measuring Measurement is the quantification of attributes of an object or event, which can be used to compare with other objects or events. In other words, measurement is a process of determining how large or small a physical quantity is as compared t ...
9–10 cm in length and weighing 7–12 grams. The adult is a compact bird with a short black tail, and stubby black and pale grey (not distinctly bluish) mandibles. It is black to brownish black on the head, chin, throat and center of the chest, with some purple-green iridescence on the face and sides of the breast. It has greyish brown upper parts and white underparts with irregular barring on the flanks and rump. A small green iridescent patch is present on the outer scapular feathers, besides, especially in the western race, the sides of the lower breast. Wing coverts and remiges are bordered in a paler or warmer tone. The sexes are similar. Immature birds are dun brown above with buff head and underpart plumage. They moult into full adult plumage by age six months, when the males also begin to sing and exhibit breeding behaviour. The race ''S. c. scutata'' has barred rather than bronzy green plumage at the junction of the lower breast and flanks. Rump and upper tail feathers only finely barred and almost uniformly dark. These characters are however somewhat variable and unpredictable. This species has a number of calls including a ' in flight, a twittering when perched, consisting of various wheezy or buzzing notes. The song is a concatenated and somewhat repetitive series of notes.


Distribution and habitat

The bronze mannikin' is native to mainland Africa and the Bioko, Pemba,
Zanzibar Zanzibar (; ; ) is an insular semi-autonomous province which united with Tanganyika in 1964 to form the United Republic of Tanzania. It is an archipelago in the Indian Ocean, off the coast of the mainland, and consists of many small islands ...
, Mafia and
Comoro islands The Comoro Islands or Comoros ( Shikomori ''Komori''; ar, جزر القمر , ''Juzur al-qamar''; french: Les Comores) form an archipelago of volcanic islands situated off the southeastern coast of Africa, to the east of Mozambique and northw ...
, and has been introduced to
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and Unincorporated ...
, where it is established. The two accepted races have an extensive region around the upper
Nile River The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest rive ...
which is inhabited by birds with intermediate features. A proposed third race, ''S. c.'' subsp. ''tressellata'' Clancey, 1964 is not generally recognized.


Behaviour and ecology

The bronze mannikin is a tiny gregarious bird which feeds mainly on seeds, including wild grass seeds,
millet Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most species generally referred to as millets belong to the tribe Paniceae, but some millets al ...
,
rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...
and
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
. Alternatively termites, nectar or strands of algae may also be eaten. Before going to roost at nightfall, they usually visit a watering hole where vegetation is hanging into the water. They roost at night in ball-shaped nests, which in the non-breeding season are built solely for this purpose. These slovenly
communal roosting Communal roosting is an animal behavior where a group of individuals, typically of the same species, congregate in an area for a few hours based on an external signal and will return to the same site with the reappearance of the signal. Environment ...
nests are dismantled (for reuse of material) and rebuilt almost daily at the same or a new location, in a communal effort. Each party, numbering 8 to 20 birds, seems to be dominated by a single adult male. The flock defends the immediate vicinity of a nest against intruders, but newcomers to a flock are easily accepted. They may associate with waxbill or other mannikin species, and may also use their vacated nests. Pairs often allopreen.


Breeding

They are incessant nest builders that may raise up to four broods a year, given favourable circumstances. The nest is a large domed grass structure in a tree, into which 4 to 8 small, white eggs are laid. Incubation takes 12 days, and chicks fledge after three weeks, and are independent in another three weeks. The chicks are reared on soft green seeds and insects.


Images

File:Lonchura cucullata 0004.jpg, A party of bronze mannikins foraging on the ground. Alarm notes of a sentry bird will warn of approaching danger File:Bronze Mannikin - Uganda H8O4101 (22366360183).jpg, An adult bronze mannikin in Uganda carries the seed head of green
Guinea grass Guinea Grass is a village in the Orange Walk District of the nation of Belize. It is 38 metres (127 feet) above sea level. According to the 2000 census, Guinea Grass had a population of 2,510 people; by 2010 the census figures showed a populat ...
for nest-building. Green ''
Digitaria ''Digitaria'' is a genus of plants in the Poaceae, grass family native to tropical and warm temperate regions but can occur in tropical, subtropical, and cooler temperate regions as well. Common names include crabgrass, finger-grass, and fonio. ...
'' and '' Sporobolus'' grasses are also used. File:Bronze Munia (Lonchura cucullata).jpg, Immature bird


References

* ''Birds of The Gambia'' by Barlow, Wacher and Disley,
BirdLife Species Factsheet


External links

*Bronze mannikin
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q1300907 bronze mannikin Birds of Sub-Saharan Africa Birds of the Comoros Birds of Southern Africa Birds of the Gulf of Guinea bronze mannikin Birds of East Africa