Bronze-winged Courser
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The bronze-winged courser or violet-tipped courser (''Rhinoptilus chalcopterus'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the family
Glareolidae Glareolidae is a family of birds in the wader suborder Charadrii. It contains two distinct groups, the pratincoles and the coursers. The atypical Egyptian plover (''Pluvianus aegyptius''), traditionally placed in this family, is now known to ...
. This species is named for its characteristic bronze-tipped feathers that are visible during flight. It is found living throughout
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, inhabiting semi-arid
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s and
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s. This is a
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
species which mainly feeds on ground-dwelling insects. Bronze-winged coursers are typically solitary, only forming
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
pairs for breeding. A female may produce 2-3 eggs per
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
, and the chicks receive parental care from both sexes when young. The bronze-winged courser is considered of Least Concern for conservation status, and is thought to be a very stable species.


Description

The bronze-winged courser is a species of small
wading birds 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
with long legs and short wings. The bronze-winged courser is the largest of the
courser The coursers are a group of birds which together with the pratincoles make up the family Glareolidae. They have long legs, short wings and long pointed bills which curve downwards. Their most unusual feature for birds classed as waders is that ...
s, with a body length of 25-29 centimeters, a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
up to 58 centimeters, and body weight between 91 and 220 grams.


Adult Recognition

The bronze-winged courser can be recognized by its characteristic head markings and overall coloration. The adult
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
differs from the juvenile plumage in this species. Mature adults have a dull grey-brown upper breast and back. The upper tail is white, while the remainder of the tail is dark brown and bordered with white. The breast features a thin black band that separates the brownish-grey upper chest from the buffy lower breast and belly. The head has distinct cream-white
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
, forecrown, and posterior eyeline, as well as buffy upper neck and throat. The facial lores and ear-coverts are darker brown-black. The underwing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
are white, contrasting with the dark black
primaries Primary elections, or direct primary are a voting process by which voters can indicate their preference for their party's candidate, or a candidate in general, in an upcoming general election, local election, or by-election. Depending on the c ...
which are tipped with
iridescent Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear to gradually change color as the angle of view or the angle of illumination changes. Examples of iridescence include soap bubbles, feathers, butterfl ...
copper to violet coloration. These copper wingtips are visible exclusively during flight–denoting the bird's common name the "bronze-winged courser". The
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is long and curved slightly downward, and appears black with a reddish-purple base. The eyes are large on the head, featuring a bright reddish-purple eye ring. The long legs are red in color. The adult bronze-winged courser in flight may be visually confused with another group that is similar in appearance–the
lapwing Lapwings (subfamily Vanellinae) are any of various ground-nesting birds (family Charadriidae) akin to plovers and dotterels. They range from in length, and are noted for their slow, irregular wingbeats in flight and a shrill, wailing cry. A gro ...
s. Both have comparable underpart coloration and body size.


Juvenile Recognition

The juvenile bronze-winged courser sports a less vibrant plumage pattern in contrast to the adults. The upper feathers of the body are cream-yellow at the ends, and the breast features an even thinner dark band. Chicks of this species are speckled in appearance, looking almost like sandy burnt vegetation.


Taxonomy

The bronze-winged courser (''Rhinoptilus chalcopterus'') is a species of courser within the family
Glareolidae Glareolidae is a family of birds in the wader suborder Charadrii. It contains two distinct groups, the pratincoles and the coursers. The atypical Egyptian plover (''Pluvianus aegyptius''), traditionally placed in this family, is now known to ...
. The coursers are a part of the order
Charadriiformes Charadriiformes (, from ''Charadrius'', the type genus of family Charadriidae) is a diverse order of small to medium-large birds. It includes about 390 species and has members in all parts of the world. Most charadriiform birds live near water an ...
in the class
Aves Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
. This species has no classified subspecies.


Range

The bronze-winged courser is found in parts of
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, and can be found residing in different areas of the continent according to season. It is thought that the bronze-winged courser spends its breeding season in the southern parts of its range, and migrates north after breeding. The breeding season spans from July to December. Historical records show the bronze-winged courser present in many localities at various times through the year, including but not limited to:
Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopia ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
,
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
,
Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...
,
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
,
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeast ...
,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label=Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalahar ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most cent ...
,
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
,
Gambia The Gambia,, ff, Gammbi, ar, غامبيا officially the Republic of The Gambia, is a country in West Africa. It is the smallest country within mainland AfricaHoare, Ben. (2002) ''The Kingfisher A-Z Encyclopedia'', Kingfisher Publicatio ...
, and more recently
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
.


Habitat and Disruption

The bronze-winged courser inhabits
semi-arid A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
environments such as dry
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s with areas of dense brush. This species can rarely be found in open areas including
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
s located nearby
wooded A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
savannas. They tend to inhabit lower elevations, though can be found within areas of elevation up to 2,350 meters. Breeding is thought to occur within a lower elevation range, likely not surpassing elevations of more than 1700 meters. The bronze-winged courser is susceptible to disturbance by human activities, particularly related to land alteration of its habitat due to farming or agricultural practices. Clearing woodlands and similar environment reduced the habitat available for bronze-winged courser. This species is also predated on by other birds in its environment, including the
Bateleur The bateleur (; ''Terathopius ecaudatus'') is a medium-sized eagle in the family Accipitridae. It is often considered a relative of the snake eagles and, like them, it is classified within the subfamily Circaetinae.Kemp, A. C., G. M. Kirwan, an ...
and the
Tawny Eagle The tawny eagle (''Aquila rapax'') is a large bird of prey. Like all eagles, it belongs to the family ''Accipitridae''. Its heavily feathered legs mark it as a member of the subfamily Aquilinae, also known as booted eagles.Helbig, A. J., Kocum, ...
.


Behavior


Vocalizations

The bronze-winged courser can be heard by its
calls Call or Calls may refer to: Arts, entertainment, and media Games * Call, a type of betting in poker * Call, in the game of contract bridge, a bid, pass, double, or redouble in the bidding stage Music and dance * Call (band), from Lahore, Paki ...
at night, but is otherwise a quiet species.


Diet

The bronze-winged courser is thought to be a strict
insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...
–presumably feeding on ground dwelling insects such as
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s. The bronze-winged courser is a nocturnal feeder, foraging in a
plover Plovers ( , ) are a widely distributed group of wading birds belonging to the subfamily Charadriinae. Description There are about 66 species in the subfamily, most of them called "plover" or "dotterel". The closely related lapwing subfa ...
-like manner–
pecking Pecking is the action of a bird using their beak to search for food or otherwise investigate an object or area by tapping it. Pecking can also be used by a bird to attack or fight another bird. Pecking is frequently observed in chickens and other ...
at the ground to capture prey while walking or running around.


Reproduction

The bronze-winged courser is a solitary species which forms
monogamous Monogamy ( ) is a form of Dyad (sociology), dyadic Intimate relationship, relationship in which an individual has only one Significant other, partner during their lifetime. Alternately, only one partner at any one time (Monogamy#Serial monogamy, ...
pairs for breeding. Though forming pairs, the couples are territorial, and keep their adjacent nests at least 100 meters from one another. Breeding occurs during the
dry season The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
. Bronze-winged coursers typically select nesting areas that have been impacted by recent burns or fires. They usually do not construct a true
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
, but rather clear a small "nest" like spot, often no more than a 1 centimeter deep indentation in the ground. The bronze-winged courser nests in open sites with good visibility. Within the nest, the bronze-winged courser lays a
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
of up to 3 eggs, which are
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
d to blend into their environments. This camouflaging of the eggs allows higher survival rates with such an open nesting environment. Egg camouflaging can vary in color, and closely mimics the environment in which they are laid. Eggs can appear cream to yellow, with hints of grey, sepia, reddish-brown, darker black, and even purple tones. Eggs are small in size, measuring anywhere from 33-40mm x 25-28mm. The eggs are incubated by both parents for 25–27 days, during which the parents alternate between sitting on the nest while the other stands watch close by. If approached by a predator, the parents will flee the nest and leave the eggs in the open. It has been found that bronze-winged coursers will move at least 5 meters away from the nest if threatened. Once hatched, the chicks receive
parental care Parental care is a behavioural and evolutionary strategy adopted by some animals, involving a parental investment being made to the evolutionary fitness of offspring. Patterns of parental care are widespread and highly diverse across the animal k ...
from both sexes. It is unknown how long the chicks
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
period lasts. The plumage on chicks of the bronze-winged courser is white from below, and appears speckled with red-brown and black from above. The legs are dark grey and the bill is black.


Status and Conservation

The bronze-winged courser is considered of
Least Concern (LC) A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
in nature protection. This species is not globally threatened, and is highly protected by the establishment of
nature Nature, in the broadest sense, is the physics, physical world or universe. "Nature" can refer to the phenomenon, phenomena of the physical world, and also to life in general. The study of nature is a large, if not the only, part of science. ...
and
game reserve A game reserve (also known as a wildlife preserve or a game park) is a large area of land where wild animals live safely or are hunted in a controlled way for sport. If hunting is prohibited, a game reserve may be considered a nature reserve; ...
s within their range. It is difficult to record true population numbers of bronze-winged coursers. This species is often underestimated and size is challenging to predict. Causes of mortality in the bronze-winged courser are often linked to traffic deaths. This nocturnal species can be found running on roads at night, where it may occasionally face fatalities from motor vehicle encounters. Other mortalities are normally due to predation by local wildlife.


References


External links

* Bronze-winged courser
Species text in The Atlas of Southern African Birds
*Bronze-winged courser
Birds of the World
*Bronze-winged courser
The Cornell Lab of Ornithology
*''Rhinoptilus chalcopterus'' (Bronze-winged courser)

{{Taxonbar, from=Q1271872
bronze-winged courser The bronze-winged courser or violet-tipped courser (''Rhinoptilus chalcopterus'') is a species of bird in the family Glareolidae. This species is named for its characteristic bronze-tipped feathers that are visible during flight. It is found livi ...
Birds of Sub-Saharan Africa
bronze-winged courser The bronze-winged courser or violet-tipped courser (''Rhinoptilus chalcopterus'') is a species of bird in the family Glareolidae. This species is named for its characteristic bronze-tipped feathers that are visible during flight. It is found livi ...
Taxonomy articles created by Polbot Taxa named by Coenraad Jacob Temminck