Bromus (section)
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''Bromus'' is a large genus of grasses, classified in its own tribe Bromeae. They are commonly known as bromes, brome grasses, cheat grasses or chess grasses. Estimates in the scientific literature of the number of species have ranged from 100 to 400, but plant taxonomists currently recognize around 160–170 species. ''Bromus'' is part of the cool-season grass lineage (subfamily Pooideae), which includes about 3300 species. Within Pooideae, ''Bromus'' is classified in tribe Bromeae (it is the only genus in the tribe). ''Bromus'' is closely related to the wheat-grass lineage (tribe Triticeae) that includes such economically important genera as ''
Triticum Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologica ...
'' (wheat), ''
Hordeum ''Hordeum'' is a genus of annual and perennial plants in the grass family. They are native throughout the temperate regions of Africa, Eurasia, and the Americas. One species, ''Hordeum vulgare'' (barley), has become of major commercial importan ...
'' (barley) and ''
Secale ''Secale'' is a genus of the grass tribe Triticeae, which is related to barley (''Hordeum'') and wheat (''Triticum''). The genus includes cultivated species such as rye (''Secale cereale'') as well as weedy and wild rye species. The most well kn ...
'' (rye).


Etymology

The generic name ''Bromus'' is derived from the Latin ''bromos'', a borrowed word from the Ancient Greek (). and mean ''oats'', but seems to have referred specifically to '' Avena sativa'' ( Hippocrates ''On Regimen in Acute Diseases'' 2.43,
Dioscorides Medicus Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of ''De materia medica'' (, On Medical Material) —a 5-vol ...
2.94, ''
Polemo Historicus Polemon (or Polemo) is the name of eminent ancient Greeks: Philosophers *Polemon (scholarch), the head of the Platonic Academy from 314–269 BC *Polemon of Athens, a 2nd-century BC Stoic philosopher, also referred to as Polemon of Ilium *Polemon o ...
'' 88) and ''
Avena barbata ''Avena barbata'' is a species of wild oat known by the common name slender wild oat. It has edible seeds. It is a diploidized autotetraploid grass (2n=4x=28). Its diploid ancestors are ''A. hirtula'' Lag. and ''A. wiestii'' Steud (2n=2x=14), whi ...
'' (Theophrastus ''
Historia Plantarum Historia may refer to: * Historia, the local version of the History channel in Spain and Portugal * Historia (TV channel), a Canadian French language specialty channel * Historia (newspaper), a French monthly newspaper devoted to History topics * ...
'' 8.9.2, Pseudo-Dioscorides 4.137). The generic name comes from , a native Latin word for "oats" or "wild oats".


Description

''Bromus'' species occur in many habitats in temperate regions of the world, including Africa,
America The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
,
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and Eurasia. There are considerable morphological differences between some species, while the morphological differences between others (usually those species that are closely related) are subtle and difficult to distinguish. As such, the taxonomy of the genus is complicated. ''Bromus'' is distinguished from other grass genera by a combination of several morphological characteristics, including leaf sheaths that are closed (connate) for most of their length, awns that are usually inserted subapically, and hairy appendages on the
ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...
. The leaf blades and sheaths, which comprise the
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
can be hairless, sparsely hairy or hairy. The inflorescence is a dense or open
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
, usually drooping or nodding, sometimes spreading (as in Japanese brome, ''B. japonicus'').


Ecology

The
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
use ''Bromus'' as a foodplant, such as the chequered skipper (''Carterocephalus palaemon'').


Uses

''Bromus'' species are generally considered to have little economic value to humans, at least in present times. ''Bromus mango'' was historically cultivated in Chile and Argentina by indigenous peoples, who used it both as fodder and food. The Tarahumara Indians in northern Mexico use the grains of some native ''Bromus'' species to aid fermentation in making one of their cultural beverages. As names like poverty brome ( ''B. sterilis'') and
ripgut brome Ripgut brome or ripgut grass refers to some species of brome grasses ('' Bromus''): * '' Bromus diandrus'' (great brome) * '' Bromus rigidus'' (stiff brome) The name refers to the fact that even for brome grasses, these species are heavily scler ...
(''B. diandrus'') attest, some species are not very useful as
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food g ...
because their leaves sclerotize quickly and may even be harmful to livestock due to the high silica content. Others, such as meadow brome ('' Bromus riparius''), native to parts of Russia, are planted as forage in the Great Plains of North America. Brome grasses are not usually grown as ornamental plants due to most species' nondescript appearance. Some are useful to prevent erosion but such use must be cautiously controlled as most ''Bromus'' have the ability to spread, becoming
invasive weed An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
s. Cheatgrass (''
Bromus tectorum ''Bromus tectorum'', known as downy brome, drooping brome or cheatgrass, is a winter annual grass native to Europe, southwestern Asia, and northern Africa, but has become invasive in many other areas. It now is present in most of Europe, southe ...
'') is a particularly troublesome weed across much of western North America (from southern British Columbia to California.)


Taxonomy and systematics

Taxonomists have generated various classification schemes to reflect the morphological variation that is seen in ''Bromus''. In North America, five morphologically similar groups of species, called sections, are generally recognized: '' Bromus'', ''
Genea ''Genea'' is a genus of truffle-like fungi in the family Pyronemataceae. There are about 32 species in the genus that occur in North America and Europe. The genus was circumscribed by Italian mycologist Carlo Vittadini in 1831. The genus na ...
'', '' Ceratochloa'', '' Neobromus'', and '' Bromopsis''. Sections ''Bromus'' and ''Genea'' are native to the Old World (Eurasia), but many species are introduced into North America. Sections ''Bromopsis'', ''Neobromus'', and ''Ceratochloa'' have several native species in North America.


Selected species


Gallery

Bromus inermis detail.jpeg,
awnless brome ''Bromus inermis'' is a species of the true grass family ( Poaceae). This rhizomatous grass is native to Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the ...
(''B. inermis'') flowers Bromus kalmii EPA-3.jpg, Arctic brome
''B. kalmii'' Bromus madritensis rubens in desert.jpg,
red brome ''Bromus madritensis'' is a species of brome grass known by the common name compact brome. The specific epithet ''madritensis'' refers to Madrid, Spain. It has a diploid number of 28. There are two subspecies: *''Bromus madritensis'' subsp. ''ma ...

''B madritensis'' ssp. ''rubens'' Bromus sitchensis 2689.JPG, Alaska brome
''B. rigidus'' Bromus secalinus2.jpg,
rye brome ''Bromus secalinus'' is a species of bromegrass known as rye brome. The specific epithet ''secalinus'' is Latin, meaning "rye-like". The fruits are hard, rounded glumes that appear superficially similar to the rye grain, which gives the brome i ...
(''B. secalinus'') flowers Bromus sterilis.jpeg,
barren brome ''Bromus sterilis'' is an Annual plant, annual or Biennial plant, biennial species of Bromus, bromegrass known as barren brome, poverty brome, and sterile brome. Description This is an Annual plant, annual or Biennial plant, biennial grass ran ...

''B. sterilis''


See also

* List of Poaceae genera


References


External links


Brome grasses UK


* {{Taxonbar, from1=Q147621, from2=Q4895613 Poaceae genera Grasses of Africa Grasses of Asia Grasses of Europe Grasses of North America Grasses of Oceania Grasses of South America