''Plantago major'', the broadleaf plantain, white man's footprint, waybread, or greater plantain, is a species of

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the plantain family

Plantaginaceae Plantaginaceae, the plantain family, is a large, diverse family of flowering plants in the order Lamiales that includes common flowers such as snapdragon and foxglove. It is unrelated to the banana-like fruit also called "plantain." In older cl ...

. The plant is native to Eurasia. The young, tender leaves can be eaten raw, and the older, stringier leaves can be boiled in stews and eaten.

Description

''Plantago major'' is a

herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...

perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...

with a rosette of

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

in diameter.Blamey, M. & Grey-Wilson, C. (1989). ''Flora of Britain and Northern Europe''. Each leaf is oval-shaped, long and broad, rarely up to long and broad, with an acute apex, a smooth margin, and a distinct petiole almost as long as the leaf itself. There are five to nine conspicuous veins over the length of the leaf. The

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s are small, greenish-brown with purple

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s, produced in a dense spike long on top of a stem tall and rarely to tall. Plantain is

wind-pollinated Anemophily or wind pollination is a form of pollination whereby pollen is distributed by wind. Almost all gymnosperms are anemophilous, as are many plants in the order Poales, including grasses, sedges, and rushes. Other common anemophilous plan ...

and propagates primarily by seeds, which are held on the long, narrow spikes which rise well above the foliage. Each plant can produce up to 20,000 seeds, which are very small and oval-shaped, with a bitter taste.

Taxonomy

Broadleaf plantain is not closely related to the fruit also known as

plantain Plantain may refer to: Plants and fruits * Cooking banana, banana cultivars in the genus ''Musa'' whose fruits are generally used in cooking ** True plantains, a group of cultivars of the genus ''Musa'' * ''Plantaginaceae'', a family of flowerin ...

, which is a kind of

banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...

. There are three

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

: *''Plantago major'' subsp. ''major''. *''Plantago major'' subsp. ''intermedia'' (DC.) Arcang. *''Plantago major'' subsp. ''winteri'' (Wirtg.) W.Ludw.

Distribution and habitat

The plant is native to most of

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

and northern and central

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

,Natural History Museum
''Plantago major''
/ref>''Flora Europaea''
''Plantago major''
/ref>''Flora of Pakistan''
''Plantago major''
/ref> but has widely

naturalised Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the i ...

elsewhere in the world.Joint Nature Conservation Committee
Greater Plantain ''Plantago major'' Linnaeus
/ref>Botanical Society of Britain and Irelan

''Plantago major'' grows in lawns and fields, along roadsides, and in other areas that have been disturbed by humans. It does particularly well in compacted or disturbed soils. It is believed to be one of the first plants to reach

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

after European colonisation. Reportedly brought to the Americas by Puritan colonizers, plantain was known among some Native American peoples by the common name "white man's footprint", because it thrived in the disturbed and damaged ecosystems surrounding European settlements. The ability of plantain to survive frequent trampling and colonize compacted soils makes it important for

soil rehabilitation Soil regeneration, as a particular form of ecological regeneration within the field of restoration ecology, is creating new soil and rejuvenating soil health by: minimizing the loss of topsoil, retaining more carbon than is depleted, boosting biodi ...

. Its roots break up

hardpan In soil science, agriculture and gardening, hardpan or soil pan is a dense layer of soil, usually found below the uppermost topsoil layer. There are different types of hardpan, all sharing the general characteristic of being a distinct soil layer ...

surfaces, while simultaneously holding together the soil to prevent

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

. The seeds of plantain are a common contaminant in

cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...

grain and other crop seeds. As a result, it now has a worldwide distribution.

Uses

The mature plant contains pliable and tough

fibres Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...

that can be used in survival situations to make small cords,

fishing line A fishing line is a flexible, high-tensile cord used in angling to tether and pull in fish, in conjunction with at least one hook. Fishing lines are usually pulled by and stored in a reel, but can also be retrieved by hand, with a fixed attachm ...

, sutures, or

braiding A braid (also referred to as a plait) is a complex structure or pattern formed by interlacing two or more strands of flexible material such as textile yarns, wire, or hair. The simplest and most common version is a flat, solid, three-strande ...

. Some

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

s are planted as

ornamentals Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate both ...

s, including 'Rubrifolia' with purple leaves, and 'Variegata' with

variegated Variegation is the appearance of differently coloured zones in the leaves and sometimes the stems and fruit of plants. Species with variegated individuals are sometimes found in the understory of tropical rainforests, and this habitat is the s ...

leaves.Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''.

Edibility

The leaves are edible as a

salad green Leaf vegetables, also called leafy greens, pot herbs, vegetable greens, or simply greens, are plant leaves eaten as a vegetable, sometimes accompanied by tender petioles and shoots. Leaf vegetables eaten raw in a salad can be called salad gre ...

when young and tender, but they quickly become tough and fibrous as they get older. The older leaves can be cooked in stews. The leaves contain

calcium Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to ...

and other minerals, and of plantain contain approximately the same amount of beta-carotene as a large

carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in color, though purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild carrot, ''Daucus carota'', nat ...

. The seeds are so small that they are tedious to gather, but they can be ground into a flour substitute or extender.

Herbal medicine

Plantain contains

phytochemical Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poisons ...

s including

allantoin Allantoin is a chemical compound with formula C4H6N4O3. It is also called 5-ureidohydantoin or glyoxyldiureide. It is a diureide of glyoxylic acid. Allantoin is a major metabolic intermediate in most organisms including animals, plants and bacter ...

aucubin Aucubin is an iridoid glycoside. Iridoids are commonly found in plants and function as defensive compounds. Iridoids decrease the growth rates of many generalist herbivores. Natural occurrences Aucubin, as other iridoids, is found in asterids ...

ursolic acid Ursolic acid (sometimes referred to as urson, prunol, malol, or 3β-hydroxyurs-12-en-28-oic acid), is a pentacyclic triterpenoid identified in the epicuticular waxes of apples as early as 1920 and widely found in the peels of fruits, as well ...

flavonoids Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...

, and asperuloside. Plantain extract has been studied for its potential health effects. Plantain leaves were used commonly in

folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...

for skin

poultice A poultice, also called a cataplasm, is a soft moist mass, often heated and medicated, that is spread on cloth and placed over the skin to treat an aching, inflamed, or painful part of the body. It can be used on wounds, such as cuts. 'Poultice' ...

s on wounds, sores, or insect stings. The root was used for fever and respiratory infections.

Gallery

References

External links

* {{Taxonbar, from=Q157154 Flora of the Dominican Republic

major Major (commandant in certain jurisdictions) is a military rank of commissioned officer status, with corresponding ranks existing in many military forces throughout the world. When used unhyphenated and in conjunction with no other indicators ...

Flora of Europe Demulcents Medicinal plants Plants described in 1753 Taxa named by Carl Linnaeus Leaf vegetables