Brisingidae
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The Brisingidae are a family of
starfish Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish ...
found only in the deep sea. They inhabit both the Atlantic and Pacific Oceans at abyssal depths, and also occur in the Southern Ocean and around Antarctica at slightly shallower depths. The family was named after
Brísingamen In Norse mythology, ''Brísingamen'' (or ''Brísinga men'') is the torc or necklace of the goddess Freyja. The name is an Old Norse compound ''brísinga-men'' whose second element is ''men'' "(ornamental) neck-ring (of precious metal), torc". The ...
, a necklace belonging to Freya from
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern period ...
that was stolen by
Loki Loki is a god in Norse mythology. According to some sources, Loki is the son of Fárbauti (a jötunn) and Laufey (mentioned as a goddess), and the brother of Helblindi and Býleistr. Loki is married to Sigyn and they have two sons, Narfi or Na ...
and hidden in the sea.


Characteristics

Members of this family have a small, Ophiurida-like disc, clearly demarcated from the arms, which number more than five. The disc is approximately circular with a rim of fused plates which gives rigidity. The madreporite is near the margin of the disc. The arms are long and tapering with the ratio of the arm length to the disc radius being greater than 6/1. There is an acute angle between them and they can be shed at the base where they are separated from the disc by a deep groove. After a narrow cylindrical portion, the part of the arms closest to the disc accommodates the
gonad A gonad, sex gland, or reproductive gland is a mixed gland that produces the gametes and sex hormones of an organism. Female reproductive cells are egg cells, and male reproductive cells are sperm. The male gonad, the testicle, produces sper ...
al tissues and widens out somewhat. Beyond this area, the arms taper to a long point. The plates on the aboral (upper) side of the arms are small, and are much smaller than the marginal plates, which are armed with spines. The ambulacral groove on the underside of each arm is broad and there is a row of tube feet with suckers on either side of the groove. The pedicellariae are crossed.


Biology

Members of this family live on hard surfaces at depths between . They raise their arms vertically above their discs to filter feed on suspended organic particles drifting past. They are able to raise their arms in this way because of the small size of its plates on the aboral surfaces of their arms which gives them flexibility in a vertical plane. '' Novodinia'' is less adept at raising its arms, because of a different arrangement of the plates; in contrast, members of the other brisingid family, Freyellidae, have not been detected raising their arms in this way.


Genera and species

Brisingidae contains the following
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
and species: *'' Astrolirus'' Fisher, 1917 **''
Astrolirus panamensis ''Astrolirus'' is a genus of echinoderms belonging to the family Brisingidae. They are found in benthic habitats in the Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the ...
'' (Ludwig, 1905) **''
Astrolirus patricki ''Astrolirus patricki'' is a species of starfish in the family Brisingidae. It is a deep-sea species found on seamounts in the northwestern Pacific Ocean, at a depth of between . This species was discovered to science in 2013, and described in ...
'' Zhang, Zhou, Xiao & Wang, 2020 *'' Astrostephane'' Fisher, 1917 ** '' Astrostephane acanthogenys'' (Fisher, 1916) ** '' Astrostephane moluccana'' (Fisher, 1916) *'' Brisinga'' Asbjørnsen, 1856 ** ''
Brisinga alberti Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** ''Brisinga analoga'' (Fisher, 1919) ** ...
'' Fisher, 1907 ** ''
Brisinga analoga Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' (Fisher, 1919) ** ''
Brisinga andamanica Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **''Brisinga alberti'' Fisher, 1907 ** ''Brisinga analoga'' (Fisher, 1919) ** ...
'' Wood-Mason & Alcock, 1891 ** ''
Brisinga bengalensis Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **''Brisinga alberti'' Fisher, 1907 ** ''Brisinga analoga'' (Fisher, 1919) ** ...
'' Wood-Mason & Alcock, 1891 ** ''
Brisinga chathamica Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' McKnight, 1973 ** ''
Brisinga costata Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Verrill, 1884 ** ''
Brisinga cricophora Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Sladen, 1889 ** ''
Brisinga distincta Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Sladen, 1889 ** '' Brisinga endecacnemos'' Asbjørnsen, 1856 ** ''
Brisinga eucoryne Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **''Brisinga alberti'' Fisher, 1907 ** ''Brisinga analoga'' (Fisher, 1919) ** ...
'' Fisher, 1916 ** ''
Brisinga evermanni Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Fisher, 1906 ** ''
Brisinga gunnii Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Alcock, 1893 ** ''
Brisinga hirsuta Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Perrier, 1894 ** ''
Brisinga insularum Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Wood-Mason & Alcock, 1891 ** ''
Brisinga panopla Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Fisher, 1906 ** ''
Brisinga parallela Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Koehler, 1909 ** ''
Brisinga synaptoma ''Brisinga synaptoma'' is a species of starfish in the family Brisingidae. It has 12 to 15 arms and a very long genital area crossed by 35 to 40 irregular costae. The scientific name of the species, as ''Craterobrisinga synaptoma'', was publishe ...
'' (Fisher, 1917) ** ''
Brisinga tasmani ''Brisinga synaptoma'' is a species of starfish in the family Brisingidae, found in deep sea waters off the coast of New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two ma ...
'' H.E.S. Clark, 1970 ** ''
Brisinga trachydisca Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **'' Brisinga alberti'' Fisher, 1907 ** '' Brisinga analoga'' (Fisher, 1919) * ...
'' Fisher, 1916 ** ''
Brisinga variispina Brisinga is a genus of starfish in the family Brisingidae. The species in this genus are primarily found in deep sea habitats. Species *''Brisinga'' Asbjørnsen, 1856 **''Brisinga alberti'' Fisher, 1907 ** ''Brisinga analoga'' (Fisher, 1919) ** ...
'' Ludwig, 1905 *'' Brisingaster'' Loriol, 1883 ** '' Brisingaster robillardi'' de Loriol, 1883 *'' Brisingella'' Fisher, 1917 ** '' Brisingella verticillata'' (Sladen, 1889) *'' Brisingenes'' Fisher, 1917 ** '' Brisingenes anchista'' Fisher, 1919 ** '' Brisingenes mimica'' (Fisher, 1916) ** '' Brisingenes multicostata'' (Verrill, 1894) ** '' Brisingenes plurispinula'' Aziz & Jangoux, 1985 *'' Hymenodiscus'' Perrier, 1884 ** '' Hymenodiscus agassizi'' (Perrier, 1882) ** '' Hymenodiscus aotearoa'' (McKnight, 1973) ** '' Hymenodiscus armillata'' (Sladen, 1889) ** '' Hymenodiscus beringiana'' (Korovchinsky, 1967) ** '' Hymenodiscus coronata'' (Sars G.O., 1872) ** '' Hymenodiscus distincta'' (Sladen, 1889) ** '' Hymenodiscus exilis'' (Fisher, 1905) ** '' Hymenodiscus fragilis'' (Fisher, 1906) ** '' Hymenodiscus membranacea'' (Sladen, 1889) ** '' Hymenodiscus monacantha'' H.L. Clark, 1920 ** '' Hymenodiscus ochotensis'' Djakonov 1950 ** '' Hymenodiscus pannychia'' Fisher, 1928 ** '' Hymenodiscus pusilla'' Fisher, 1917 ** '' Hymenodiscus submembranacea'' (Doderlein, 1927) ** '' Hymenodiscus tenella'' (Luwig, 1905) ** '' Hymenodiscus verticellata'' (Sladen, 1889) *''
Labidiaster ''Labidiaster'' is a genus of echinoderms belonging to the family Heliasteridae. The species of this genus are found in the coasts of Antarctica and southernmost South America. Species Two species are recognized: *''Labidiaster annulatus ''L ...
'' Lütken, 1872 *'' Midgardia'' Downey, 1972 ** '' Midgardia xandaros'' Downey, 1972 *'' Novodinia'' Dartnall, Pawson, Pope & B.J. Smith, 1969 ** ''
Novodinia americana Novodina americana is a species of starfish found of the coast of Greenland. A 2018 study of deep-water starfish species found that ''N. americana'' was both bioluminescent and, unexpectedly for an inhabitant of the aphotic The aphotic zone (a ...
'' (Verrill, 1880) ** ''
Novodinia antillensis ''Novodinia antillensis'', the velcro sea star, is a species of starfish in the family Brisingidae. It is found in the deep sea in the tropical and subtropical western Atlantic Ocean, being quite common at a depth of around on the Mesoamerican ...
'' (A.H.Clark, 1934) ** ''
Novodinia austini ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *''Novodinia australis'' *'' ...
'' (Koehler, 1909) ** ''
Novodinia australis ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (H.L. Clark, 1916) ** ''
Novodinia clarki ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (Koehler, 1909) ** ''
Novodinia homonyma ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' Downey, 1986 ** ''
Novodinia magister ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' Fisher, 1917 ** ''
Novodinia novaezelandiae ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (H.E.S. Clark, 1962) ** ''
Novodinia pacifica ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis ''Novodinia antillensis'', the velcro sea star, is a s ...
'' (Fisher, 1906) ** ''
Novodinia pandina ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (Sladen, 1889) ** ''
Novodinia penichra ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (Fisher, 1916) ** ''
Novodinia radiata ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' Aziz & Jangoux, 1985 ** ''
Novodinia semicoronata ''Novodinia'' is a genus of echinoderms belonging to the family Brisingidae. The genus has almost cosmopolitan distribution. Species: *'' Novodinia americana'' *''Novodinia antillensis'' *'' Novodinia austini'' *'' Novodinia australis'' *' ...
'' (Perrier, 1885) *'' Odinella'' Fisher, 1940 **'' Odinella nutrix'' Fisher, 1940 *'' Stegnobrisinga'' Fisher, 1916 **'' Stegnobrisinga gracilis'' (Koehler, 1909) **'' Stegnobrisinga placoderma'' Fisher, 1916 **'' Stegnobrisinga splendens'' H.L. Clark, 1926


References

{{Taxonbar, from=Q3466573 Brisingida Echinoderm families