Breviparopus
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''Breviparopus'' (ichnotype ''B. taghbaloutensis'') is the name given to an ichnogenus of
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
, having been made by an unknown
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
. As an ichnogenus, the taxon is represented by (and named for) a 90-metre (295 ft) long series of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
tracks, or
ichnite A fossil track or ichnite (Greek "''ιχνιον''" (''ichnion'') – a track, trace or footstep) is a fossilized footprint. This is a type of trace fossil. A fossil trackway is a sequence of fossil tracks left by a single organism. Over the year ...
s, found in the spring of 1979 in the
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through Moroc ...
of present-day
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
. At the time, this area would have been part of the splitting
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
supercontinent. The animal that produced the ''Breviparopus'' tracks is rumored to be one of the largest dinosaurs, though its exact size has been the subject of much debate.Wedel, M. (2009).
How big were the biggest sauropod trackmakers?
''Sauropod Vertebra Picture of the Week'', 13 October 2009, updated 3 December 2009. Accessed 1 November 2010.


Fossil material and age

The combined length of the tracks of the foot and hand measured , and they were in width. They are commonly dated to the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
period (approximately 160–175 million years ago), though they are more likely from the early Cretaceous, roughly 130-120 million years ago. They were first described by Jean-Michel Dutuit and Achmed Ouazzou in 1980.


Size of trackmaker

The size, weight, and even
family tree A family tree, also called a genealogy or a pedigree chart, is a chart representing family relationships in a conventional tree structure. More detailed family trees, used in medicine and social work, are known as genograms. Representations of ...
of the animal that made ''Breviparopus'' tracks is unknown, but a great deal of speculation has arisen about this animal. Length estimates as great as have been given in popular books such as ''
Guinness World Records ''Guinness World Records'', known from its inception in 1955 until 1999 as ''The Guinness Book of Records'' and in previous United States editions as ''The Guinness Book of World Records'', is a reference book published annually, listing world ...
'', though these were based on the misconception that the figure was based on a single footprint, not the combined length of the fore and hind feet. In reality, the trackmakers of ''Breviparopus'' were likely somewhat smaller than this. The actual prints are wideIshigaki, S. (1989) ''Footprints of swimming sauropods from Morocco''; pp. 83-86 in Gillette, D. D. and Lockley, M. G. (eds.), ''Dinosaur Tracks and Traces''. Cambridge University Press, Cambridge. and about as long. The feet were probably even larger than wide since the edges of the prints are collapsed. This would result in an animal roughly long, assuming proportions similar to '' Giraffatitan''. Earlier estimates that the prints were only wide, no larger than the feet of ''
Diplodocus ''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπ ...
'', have since been shown to be incorrect. In 2020 Molina-Pérez and Larramendi suggested that the narrow-gauge track, the position of the claws, and the era all indicate that it belonged to an enormous diplodocoid, and estimated its size at and .


Taxonomy

Traditionally, ''Breviparopus'' has been regarded as a
brachiosaur The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the le ...
, which seemed to correspond with the fact that ''
"Brachiosaurus" nougaredi ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in t ...
'', a brachiosaur known from a colossal
sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
, was discovered not too far from Morocco, in Early Cretaceous deposits in Wargla, Algeria. Since no actual bones of ''Breviparopus'' have been identified, it is difficult to say with certainty whether the tracks were made by a
diplodocid Diplodocids, or members of the family Diplodocidae ("double beams"), are a group of sauropod dinosaurs. The family includes some of the longest creatures ever to walk the Earth, including ''Diplodocus'' and ''Supersaurus'', some of which may have ...
,
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still th ...
or
brachiosaur The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the le ...
. All that can be said for certain, according to Michel Monbaron and colleagues, is that ''Breviparopus'' is quite distinct from the prints made by ''
Atlasaurus ''Atlasaurus'' is a genus of sauropod dinosaurs from Middle Jurassic (Bathonian to Callovian stages) beds in North Africa.M. Monbaron, D. A. Russell, and P. Taquet. (1999). ''Atlasaurus imelakei'' n.g., n.sp., a brachiosaurid-like sauropod from t ...
''. However, the presence of a small medially directed thumb-claw print makes it likely the animal was a brachiosaur, since they have small thumb claws at ground level (as opposed to diplodocids and camarasaurids, whose thumb metacarpals are short and hold their thumb claws off the ground), and the narrow-gauge stance also fits with a brachiosaur body shape - in the Early Cretaceous, the dominant sauropods were wide-bodied titanosaurs, and deep-bodied brachiosaurs which had more narrowly spaced legs - therefore the narrow gauge of the ''Breviparopus'' prints favors a brachiosaurid diagnosis (this seems to be corroborated by the sacrum of ''
Brachiosaurus nougaredi ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in the ...
'', which is unusually narrow for its length even by brachiosaur standards). It is also worth noting that the bone fossil record from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
, according to C.A. Meyer is rather more incomplete and fragmented than the fossil record of dinosaur tracks.Meyer, C.A. and Thüring, B. "Mind the 'Middle Jurassic' Gap - Bone versus Track Record in Dinosaurs.

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See also

* List of dinosaur ichnogenera * '' Maraapunisaurus'', another gigantic sauropod * Largest prehistoric animals * Dinosaur size


References


External links


Artist's rendering of 14 brachiosaurs, including ''Breviparopus'' scaled to the known 90 cm-wide tracks from Morocco.

First life restoration of Breviparopus by Nima Sassani, same artist as the above.
{{Taxonbar, from=Q2778303 Dinosaur trace fossils