Brachysuchus Megalodon
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''Brachysuchus'' (meaning "wide crocodile") is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of phytosaur known from the late
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
period ( Carnian stage) of Dockum Group in
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
, United States. It is known from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
UMMP 10336 is composed of a
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
, lower
jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...
s and partial
postcranium Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...
and from the associated paratype UMMP 14366, nearly complete skull, recovered from the 'Pre-Tecovas Horizon' in the Dockum Group. It was first named by Case in
1929 This year marked the end of a period known in American history as the Roaring Twenties after the Wall Street Crash of 1929 ushered in a worldwide Great Depression. In the Americas, an agreement was brokered to end the Cristero War, a Catholic ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Brachysuchus megalodon''. Its closest relative was ''
Angistorhinus ''Angistorhinus'' (meaning "narrow snout" or "hook snout") is an extinct genus of phytosaur known from the Late Triassic period of Texas and Wyoming, United States. It was first named by Mehl in 1913 and the type species is ''Angistorhinus grand ...
''. However, its rostral crest was much smaller than that of ''Angistorhinus,'' and the rostrum as a whole is shorter and thicker.


Features

The lower jaws of ''Brachysuchus'' are expanded at the tip to form a large bulge, holding the creature's largest tusks. The surface of this part of the
bone A bone is a Stiffness, rigid Organ (biology), organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red blood cell, red and white blood cells, store minerals, provid ...
has a wrinkled look, with many
blood vessel The blood vessels are the components of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away ...
s running through it. In the lower jaw, there are three tusks in each side of this protuberance. Behind this part the jaws have fused together for a little under half their length before diverging after thirty-one of forty-six post-protuberance teeth. While some of the tusks at the tip are missing, most of the rest of the teeth in the jaw are still present. There are new tusks forming in the sockets, indicating that ''Brachysuchus'' regrew teeth all its life. There is a large
foramen In anatomy and osteology, a foramen (;Entry "foramen"
in
between the dentary, the angular and the prearticular that passes right through the jaw (visible on the picture). The teeth themselves are asymmetrical in
cross-section Cross section may refer to: * Cross section (geometry) ** Cross-sectional views in architecture & engineering 3D *Cross section (geology) * Cross section (electronics) * Radar cross section, measure of detectability * Cross section (physics) **Ab ...
, with the outer side more convex than the inner side. This asymmetry increases from front to back. They are long and conical, and those from the ninth to twenty-ninth have flutings on the sides. From the thirtieth to the forty-ninth, they have no flutings but are
crenulate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
d and broaden antero-posteriorly. This suggests that these teeth were better for slicing
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
than the previous tusks and conical teeth, which would have impaled and injured it. In parts of the jaws the fused section in the middle rises so high that the teeth could only grip prey and could probably have done little damage. The prearticular is fused with the articular. The skull is around 125 cm long, with a distinctive hooked tip. It is much shorter and broader than that of many phytosaurs such as ''
Leptosuchus ''Leptosuchus'' is an extinct genus of leptosuchomorph phytosaur with a complex taxonomical history. Fossils have been found from the Dockum Group and lower Chinle Formation outcropping in Texas, New Mexico, and Arizona, USA, and date back to th ...
'' or ''Angistorhinus''. The eye sockets are narrow and long, and the
fenestrae A fenestra (fenestration; plural fenestrae or fenestrations) is any small opening or pore, commonly used as a term in the biological sciences. It is the Latin word for "window", and is used in various fields to describe a pore in an anatomical st ...
are smaller and less rounded than ''Angistorhinus.'' It has a comparatively small rostral crest. There are several rough surfaces at the posterior end denoting probable muscle attachments, which show that it had very powerful jaw muscles and a strong bite. Overall ''Brachysuchus'' was approximately eight metres long, much bigger than most contemporary phytosaurs. It had extremely heavy jaws and a relatively short fused region, giving it a wider bite. There is also evidence of heavily developed jaw muscles. This, as well as the cutting teeth mentioned earlier, indicates that ''Brachysuchus'' preyed on different animals to its thinner-jawed relatives. Instead of using its jaws to catch
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, it was able to attack and feed off the heavily armoured
stegocephalia Stegocephali (often spelled Stegocephalia) is a group containing all four-limbed vertebrates. It is equivalent to a broad definition of Tetrapoda: under this broad definition, the term "tetrapod" applies to any animal descended from the first ve ...
ns that roamed the surrounding area, and would probably have eaten other phytosaurs if food was scarce. The slightly deformed tip of one specimen's lower jaw suggests that it used a lot of force to break through its prey's armour.


References

{{Taxonbar, from=Q4953588 Phytosaurs Prehistoric reptile genera Carnian genera Late Triassic reptiles of North America Triassic geology of Texas Paleontology in Texas Fossil taxa described in 1929 Taxa named by Ermine Cowles Case