The genus ''Bowenia'' includes two

living Living or The Living may refer to: Common meanings *Life, a condition that distinguishes organisms from inorganic objects and dead organisms ** Living species, one that is not extinct *Personal life, the course of an individual human's life * ...

and two

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...

s in the family

Stangeriaceae Stangeriaceae is the smallest family of the cycads, including only two extant genera, '' Stangeria'' and ''Bowenia'', which both have an underground stem. Taxonomy Although the family was first published by Schimper and Schenk as Stangerieae, t ...

, sometimes placed in their own family Boweniaceae. They are entirely restricted to

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

. The two living species occur in

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

. ''B. spectabilis'' grows in warm, wet, tropical

rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...

s, on protected slopes and near streams, primarily in the lowlands of the Wet Tropics Bioregion. However, it has a local form with serrate pinna margins that grows in rainforest, ''Acacia''-dominated transition forest, and also ''Casuarina''-dominated sclerophyll forest on the

Atherton Tableland The Atherton Tableland is a fertile plateau which is part of the Great Dividing Range in Queensland, Australia. The principal river flowing across the plateau is the Barron River. It was dammed to form an irrigation reservoir named Lake Tina ...

, where it is subject to periodic bushfire. ''B. serrulata'' grows in

sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaf, leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is paral ...

forest and transition forest close to the

Tropic of Capricorn The Tropic of Capricorn (or the Southern Tropic) is the circle of latitude that contains the subsolar point at the December (or southern) solstice. It is thus the southernmost latitude where the Sun can be seen directly overhead. It also reac ...

Species

The fossil species ''Bowenia eocenica'' is known from deposits in a

coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal is formed when dea ...

mine in Victoria,

, and ''B. papillosa'' is known from deposits in

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

. Both fossils are of

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

age, and consist of

leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...

let fragments. Bowenia spectabilis Daintree 2

References

External links

* * *
Gymnosperm Database, ''Bowenia''
* * * {{Taxonbar, from=Q134570 Endemic flora of Australia Cycadophyta of Australia Flora of Queensland Taxa named by Joseph Dalton Hooker