Botanical Geography
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Phytogeography (from Greek φυτόν, ''phytón'' = "plant" and γεωγραφία, ''geographía'' = "geography" meaning also distribution) or botanical geography is the branch of
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
that is concerned with the geographic distribution of
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and their influence on the earth's surface. Phytogeography is concerned with all aspects of plant distribution, from the controls on the distribution of individual species ranges (at both large and small scales, see
species distribution Species distribution —or species dispersion — is the manner in which a biological taxon is spatially arranged. The geographic limits of a particular taxon's distribution is its range, often represented as shaded areas on a map. Patterns of ...
) to the factors that govern the composition of entire communities and
flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...
s. Geobotany, by contrast, focuses on the geographic space's influence on plants.


Fields

Phytogeography is part of a more general science known as
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
. Phytogeographers are concerned with patterns and process in plant distribution. Most of the major questions and kinds of approaches taken to answer such questions are held in common between phyto- and zoogeographers. Phytogeography in wider sense (or geobotany, in German literature) encompasses four fields, according with the focused aspect, environment,
flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...
( taxa),
vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
( plant community) and origin, respectively: * plant ecology (or
mesology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
– however, the physiognomic-ecological approach on
vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
and
biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
study are also generally associated with this field); * plant geography (or phytogeography in strict sense,
chorology Chorology (from Greek , ''khōros'', "place, space"; and , ''-logia'') can mean * the study of the causal relations between geographical phenomena occurring within a particular region * the study of the spatial distribution of organisms (biogeograp ...
, floristics); * plant sociology (or phytosociology, synecology – however, this field does not prescind from flora study, as its approach to study vegetation relies upon a fundamental unit, the plant association, which is defined upon flora). * historical plant geography (or
paleobotany Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...
, paleogeobotany) Phytogeography is often divided into two main branches: ecological phytogeography and
historical phytogeography History (derived ) is the systematic study and the documentation of the human activity. The time period of event before the invention of writing systems is considered prehistory. "History" is an umbrella term comprising past events as well ...
. The former investigates the role of current day biotic and abiotic interactions in influencing plant distributions; the latter are concerned with historical reconstruction of the origin, dispersal, and extinction of taxa.


Overview

The basic data elements of phytogeography are occurrence records (presence or absence of a species) with operational geographic units such as political units or geographical coordinates. These data are often used to construct phytogeographic provinces ( floristic provinces) and elements. The questions and approaches in phytogeography are largely shared with zoogeography, except zoogeography is concerned with animal distribution rather than plant distribution. The term phytogeography itself suggests a broad meaning. How the term is actually applied by practicing scientists is apparent in the way periodicals use the term. The '' American Journal of Botany'', a monthly primary research journal, frequently publishes a section titled "Systematics, Phytogeography, and Evolution." Topics covered in the ''American Journal of Botanys "Systematics and Phytogeography" section include phylogeography, distribution of genetic variation and,
historical biogeography Phylogeography is the study of the historical processes that may be responsible for the past to present geographic distributions of genealogical lineages. This is accomplished by considering the geographic distribution of individuals in light of ge ...
, and general plant species distribution patterns. Biodiversity patterns are not heavily covered. A flora is the group of all plant species in a specific period of time or area, in which each species is independent in abundance and relationships to the other species. The group or the flora can be assembled in accordance with floral element, which are based on common features. A flora element can be a genetic element, in which the group of species share similar genetic information i.e. common evolutionary origin; a migration element has a common route of access into a habitat; a historical element is similar to each other in certain past events and an ecological element is grouped based on similar environmental factors. A population is the collection of all interacting individuals of a given species, in an area. An area is the entire location where a species, an element or an entire flora can occur. Aerography studies the description of that area, chorology studies their development. The local distribution within the area as a whole, as that of a swamp shrub, is the topography of that area. Areas are an important factor is forming an image about how species interaction result in their geography. The nature of an area’s margin, their continuity, their general shape and size relative to other areas, make the study of area crucial in identifying these types of information. For example, a relict area is an area surviving from an earlier and more exclusive occurrence. Mutually exclusive plants are called vicarious (areas containing such plants are also called vicarious). The earth’s surface is divided into floristic region, each region associated with a distinctive flora.


History

Phytogeography has a long history. One of the subjects earliest proponents was Prussian naturalist
Alexander von Humboldt Friedrich Wilhelm Heinrich Alexander von Humboldt (14 September 17696 May 1859) was a German polymath, geographer, naturalist, explorer, and proponent of Romantic philosophy and science. He was the younger brother of the Prussian minister, p ...
, who is often referred to as the "father of phytogeography". Von Humboldt advocated a quantitative approach to phytogeography that has characterized modern plant geography. Gross patterns of the distribution of plants became apparent early on in the study of plant geography. For example,
Alfred Russel Wallace Alfred Russel Wallace (8 January 1823 – 7 November 1913) was a British naturalist, explorer, geographer, anthropologist, biologist and illustrator. He is best known for independently conceiving the theory of evolution through natural se ...
, co-discoverer of the principle of natural selection, discussed the latitudinal gradients in species diversity, a pattern observed in other organisms as well. Much research effort in plant geography has since then been devoted to understanding this pattern and describing it in more detail. In 1890, the United States Congress passed an act that appropriated funds to send expeditions to discover the geographic distributions of plants (and animals) in the United States. The first of these was The Death Valley Expedition, including Frederick Vernon Coville, Frederick Funston, Clinton Hart Merriam, and others. Research in plant geography has also been directed to understanding the patterns of adaptation of species to the environment. This is done chiefly by describing geographical patterns of trait/environment relationships. These patterns termed ecogeographical rules when applied to plants represent another area of phytogeography.


Floristic regions

Floristic {{Short pages monitor