Boophis Goudoti

''Boophis'' is the only genus in the mantellid frog subfamily Boophinae. They are commonly known as bright-eyed or skeleton frogs. They show typical 'tree frog' traits, and are a good example of convergent evolution with morphologically similar species in the families Hylidae and Rhacophoridae, among others. This genus can only be found on Madagascar and Mayotte Island (Comoros).

Taxonomy

The genus ''Boophis'' was described by Johann Jakob von Tschudi in 1838. It was originally considered a member of the African-Asian family Rhacophoridae, but was moved to the family Mantellidae in 2001, into its own subfamily, Boophinae Vences & Glaw, 2001.

Morphology

''Boophis'' are arboreal frogs, exhibiting many traits that define 'tree frogs', such as expanded toe discs, long hindlimbs, and large eyes. ''Boophis'' are especially characterised by bright colouration of the iris, which is typically intricately patterned towards its inside, and often green or blue, but occasionally also red, purple, or yellow in the outer iris area. This has led to the vernacular name of bright-eyed frogs for the genus. Many species of ''Boophis'' have almost translucent skin, allowing bones and internal organs to be observed as in the unrelated glass frogs (Centrolenidae) of the tropical Americas. This has led to the vernacular name skeleton frogs for some members of the genus.

Species

The genus has presently nearly 80 species; new ones are being described every few months on average. This list may not be exhaustive.

* ''Boophis albilabris'' (Boulenger, 1888)
* ''Boophis albipunctatus'' Glaw & Thiesmeier, 1993
* ''Boophis andohahela'' Andreone, Nincheri & Piazza, 1995
* ''Boophis andrangoloaka'' (Ahl, 1928)
* ''Boophis andreonei'' Glaw & Vences, 1994
* ''Boophis anjanaharibeensis'' Andreone, 1996
* ''Boophis ankarafensis'' Penny, Andreone, Crottini, Holderied, Rakotozafy, Schwitzer & Rosa, 2014
* ''Boophis ankaratra'' Andreone, 1993
* ''Boophis arcanus'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis axelmeyeri'' Vences, Andreone & Veities, 2005
* ''Boophis baetkei'' Köhler, Glaw & Vences, 2008
* ''Boophis blommersae'' Glaw & Vences, 1994
* '' Boophis boehmei'' Glaw & Vences, 1992
* '' Boophis boppa'' Hutter & Lambert, 2015
* ''Boophis bottae'' Vences & Glaw, 2002
* ''Boophis brachychir'' Boettger, 1882
* '' Boophis burgeri'' Glaw & Vences, 1994
* ''Boophis calcaratus'' Vallan, Vences & Glaw, 2010
* '' Boophis doulioti'' Angel, 1934
* ''Boophis elenae'' Andreone, 1993
* ''Boophis englaenderi'' Glaw & Vences, 1994
* ''Boophis entingae'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* '' Boophis erythrodactylus'' (Guibé, 1953)
* '' Boophis fayi'' Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
* ''Boophis feonnyala'' Glaw, Vences, Andreone & Vallan, 2001
* ''Boophis goudotii'' Tschudi, 1838
* ''Boophis guibei'' (McCarthy, 1978)
* ''Boophis haematopus'' Glaw, Vences, Andreone & Vallan, 2001
* ''Boophis haingana'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis idae'' (Steindachner, 1867)
* ''Boophis jaegeri'' Glaw & Vences, 1992 – Green Skeleton Frog
* '' Boophis laurenti'' Guibé, 1947
* '' Boophis liami'' Vallan, Vences & Glaw, 2003
* ''Boophis lichenoides'' Vallan, Glaw, Andreone & Cadle, 1998
* ''Boophis lilianae'' Köhler, Glaw & Vences, 2008
* '' Boophis luciae'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* '' Boophis luteus'' (Boulenger, 1882)
* ''Boophis madagascariensis'' (Peters, 1874)
* ''Boophis majori'' (Boulenger, 1896)
* '' Boophis mandraka'' Blommers-Schlösser, 1979
* '' Boophis marojezensis'' Glaw & Vences, 1994
* ''Boophis masoala'' Glaw, Scherz, Prötzel & Vences, 2018
* ''Boophis miadana'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis microtympanum'' (Boettger, 1881)
* ''Boophis miniatus'' (Mocquard, 1902)
* ''Boophis narinsi'' Vences, Gehara, Köhler & Glaw, 2012
* ''Boophis nauticus'' Glaw, Hawlitschek, Glaw, and Vences, 2019
* '' Boophis obscurus'' (Boettger, 1913)
* '' Boophis occidentalis'' Glaw & Vences, 1994
* '' Boophis opisthodon'' (Boulenger, 1888)
* ''Boophis pauliani'' (Guibé, 1953)
* ''Boophis periegetes'' Cadle, 1995
* ''Boophis picturatus'' Glaw, Vences, Andreone & Vallan, 2001
* ''Boophis piperatus'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis popi'' Köhler, Glaw, Rosa, Gehring, Pabijan, Andreone & Vences, 2011
* '' Boophis praedictus'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis pyrrhus'' Glaw, Vences, Andreone & Vallan, 2001
* ''Boophis quasiboehmei'' Vences, Köhler, Crottini & Glaw, 2010
* '' Boophis rappiodes'' (Ahl, 1928)
* ''Boophis reticulatus'' Blommers-Schlösser, 1979
* '' Boophis rhodoscelis'' (Boulenger, 1882)
* ''Boophis roseipalmatus'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis rufioculis'' Glaw & Vences, 1997
* ''Boophis sambirano'' Vences & Glaw, 2005
* ''Boophis sandrae'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis schuboeae'' Glaw & Vences, 2002
* ''Boophis septentrionalis'' Glaw & Vences, 1994
* ''Boophis sibilans'' Glaw & Thiesmeier, 1993
* ''Boophis solomaso'' Vallan, Vences & Glaw, 2003
* '' Boophis spinophis'' Glaw, Köhler, de la Riva, Vieites & Vences, 2010
* ''Boophis tampoka'' Köhler, Glaw & Vences, 2007
* ''Boophis tasymena'' Vences & Glaw, 2002
* ''Boophis tephraeomystax'' (Duméril, 1853)
* ''Boophis tsilomaro'' Vences, Andreone, Glos & Glaw, 2010
* ''Boophis ulftunni'' Wollenberg, Andreone, Glaw & Vences, 2008
* ''Boophis viridis'' Blommers-Schlösser, 1979
* ''Boophis vittatus'' Glaw, Vences, Andreone & Vallan, 2001
* '' Boophis williamsi'' (Guibé, 1974)
* '' Boophis xerophilus'' Glaw & Vences, 1997

References

{{Taxonbar, from=Q2322420

Amphibians of Sub-Saharan Africa
Endemic fauna of Madagascar
Amphibian genera
Fauna of Mayotte
Taxa named by Johann Jakob von Tschudi
Taxonomy articles created by Polbot