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Bookworm is a general name for any
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
that is said to bore through
book A book is a medium for recording information in the form of writing or images, typically composed of many pages (made of papyrus, parchment, vellum, or paper) bound together and protected by a cover. The technical term for this physical arr ...
s. The damage to books that is commonly attributed to "bookworms" is often caused by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of various types of insects including
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s,
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s and
cockroach Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as ...
es, which may bore or chew through books seeking food. The damage is not caused by any species of
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...
. Some such larvae exhibit a superficial resemblance to worms and are the likely inspiration for the term, though they are not true worms. In other cases,
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s,
carpenter ant Carpenter ants (''Camponotus'' spp.) are large () ants indigenous to many forested parts of the world. They build nests inside wood consisting of galleries chewed out with their mandibles or jaws, preferably in dead, damp wood. However, unlik ...
s, and
woodboring beetle The term woodboring beetle encompasses many species and families of beetles whose larval or adult forms eat and destroy wood (i.e., are xylophagous). In the woodworking industry, larval stages of some are sometimes referred to as woodworms. The ...
s will first infest wooden bookshelves and later feed on books placed upon the shelves, attracted by the wood-pulp
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distributed ...
used in most commercial book production. True book-borers are uncommon. The primary food sources for many "bookworms" are the leather or cloth bindings of a book, the glue used in the binding process, or molds and fungi that grow on or inside books. When the pages themselves are attacked, a gradual encroachment across the surface of one page or a small number of pages is typical, rather than the boring of holes through the entire book. The term has come to have a second, idiomatic use, indicative of a person who reads a great deal or to perceived excess: someone who devours books
metaphor A metaphor is a figure of speech that, for rhetorical effect, directly refers to one thing by mentioning another. It may provide (or obscure) clarity or identify hidden similarities between two different ideas. Metaphors are often compared wit ...
ically.


Booklice

The booklouse, also known as a paperlouse, is a soft-bodied, wingless insect in the order
Psocoptera Psocoptera are a paraphyletic group of insects that are commonly known as booklice, barklice or barkflies. The name Psocoptera has been replaced with Psocodea in recent literature, with the inclusion of the former order Phthiraptera into Psocode ...
(usually ''
Trogium pulsatorium ''Trogium pulsatorium'', known generally as larger pale booklouse, is a species of granary booklouse in the family Trogiidae. Other common names include the deathwatch, common booklouse, and grain psocid. It is found in Africa, Australia, Europe ...
''), typically 1 mm or less in length. Booklice feed on microscopic
mold A mold () or mould () is one of the structures certain fungus, fungi can form. The dust-like, colored appearance of molds is due to the formation of Spore#Fungi, spores containing Secondary metabolite#Fungal secondary metabolites, fungal seco ...
s and other organic matter found in or on aging items that have been stored in places that lack the climate control necessary to inhibit organic growth. Areas of archives, libraries, and museums that are cool, damp, dark, and generally undisturbed are common sites for such growth, generating a food source which subsequently attracts booklice. Booklice will also attack bindings, glue, and paper. Despite their name, booklice are not considered to be true
lice Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a result o ...
, as they do not feed on a living host. By the 20th century, bookbinding materials had developed a high resistance against damage by various types of book-boring insects. Many museums and archives in possession of materials vulnerable to booklouse damage employ pest control methods to manage existing infestations and make use of climate control to prevent the growth of potential booklouse food sources.


Other book-eating insects


Beetles

Of the quarter million species of beetles, some adults damage books by eating paper and binding materials themselves. However, their larvae do the most damage. Typically eggs are laid on the book's edges and spine. Upon hatching, they bore into, and sometimes even through, the book.


Woodboring beetles

*
Common furniture beetle The common furniture beetle or common house borer (''Anobium punctatum'') is a woodboring beetle originally from Europe but now distributed worldwide. In the larval stage it bores in wood and feeds upon it. Adult ''Anobium punctatum'' measure in ...
*
Deathwatch beetle The deathwatch beetle (''Xestobium rufovillosum'') is a species of woodboring beetle that sometimes infests the structural timbers of old buildings. The adult beetle is brown and measures on average long. Eggs are laid in dark crevices in old w ...
*The genus '' Gastrallus'' * Indian bookworm beetle * Australian spider beetle * Cigarette beetle *
Drugstore beetle The drugstore beetle (''Stegobium paniceum''), also known as the bread beetle, biscuit beetle, and misnamed as the biscuit weevil (despite not being a true weevil), is a tiny, brown beetle that can be found infesting a wide variety of dried pla ...


Auger beetles

*
Lesser grain borer ''Rhyzopertha'' is a monotypic taxon, monotypic genus of beetles in the family Bostrichidae, the false powderpost beetles. The sole species, ''Rhyzopertha dominica'', is known commonly as the lesser grain borer, American wheat weevil, Australian ...


Long horned beetles

* House longhorn beetle


Bark beetles A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family ( Curculionidae). Although t ...

* Flat bark beetle * Merchant beetle *
Sawtoothed grain beetle ''Oryzaephilus surinamensis'', the sawtoothed grain beetle, is a beetle in the superfamily ''Cucujoidea''. It is a common, worldwide pest of grain and grain products as well as chocolate, drugs, and tobacco. The species's binomial name, meaning ...


True weevils

*
Rice weevil Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera ''Zizania'' and '' Porteresia'', both wild and domesticated, ...
*
Wheat weevil The wheat weevil (''Sitophilus granarius''), also known as the grain weevil or granary weevil, is an insect that feeds on cereal grains, and is a common pest in many places. It can cause significant damage to harvested stored grains and may drasti ...


Skin beetles

These beetles have been known to feed on leather bindings. * Furniture carpet beetle * Museum beetle * Common carpet beetle *
Varied carpet beetle The varied carpet beetle (''Anthrenus verbasci'') is a 3 mm-long beetle belonging to the family Dermestidae. They are a common species, often considered a pest of domestic houses and, particularly, natural history museums, where the larvae m ...
*
Fur beetle ''Attagenus pellio'', the fur beetle or carpet beetle, is a pest which damages stored products such as furs, skins, textiles and grain. ''Attagenus pelio'' is a 4–6 mm-long oval shaped insect with two patches of white hair on the elyt ...
*
Black carpet beetle The black carpet beetle (''Attagenus unicolor'') is a beetle that can be a serious household pest. The larvae grow to in length, are reddish brown in colour and covered with bristles. The larval form feeds on natural fibres, damaging carpets, ...
*''
Dermestes coarctatus ''Dermestes'' is a genus of beetles in the family Dermestidae, the skin beetles. The genus is distributed worldwide.Magni, P. A., et al. (2015)A biological and procedural review of forensically significant ''Dermestes'' species (Coleoptera: Derm ...
'' * Larder beetle *''
Dermestes maculatus ''Dermestes maculatus'' is a species of beetle with a worldwide distribution, being present on all continents except Antarctica. In Europe, it is present in all countries. Description The adult beetle is 5.5-10 mm long, and black with a band of ...
'' *'' Dermestes vorax'' * Khapra beetle *''
Reesa ''Reesa'' is a monotypic genus of beetles in the family Dermestidae, the skin beetles. The sole species is ''Reesa vespulae''. This beetle is native to the Nearctic, but today it can be found nearly worldwide; it is easily introduced to new are ...
'' *'' Trogoderma versicolor'' * Odd beetle


Powderpost beetle Powderpost beetles are a group of seventy species of woodboring beetles classified in the insect subfamily Lyctinae. These beetles, along with spider beetles, death watch beetles, common furniture beetles, Dermestidae, skin beetles, and others ...
s

* African powderpost beetle * Brown powderpost beetle * Black powderpost beetle


Darkling beetle Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution. Taxonomy ''Tenebrio'' is the Latin gen ...
s

*
Confused flour beetle The confused flour beetle (''Tribolium confusum''), a type of darkling beetle known as a flour beetle, is a common pest insect known for attacking and infesting stored flour and grain. They are one of the most common and most destructive insect ...
* Destructive flour beetle * Dark mealworm beetle *
Mealworm Mealworms are the larval form of the yellow mealworm beetle, ''Tenebrio molitor'', a species of darkling beetle. Like all holometabolic insects, they go through four life stages: egg, larva, pupa, and adult. Larvae typically measure about or ...
*
Red flour beetle The red flour beetle (''Tribolium castaneum'') is a species of beetle in the family Tenebrionidae, the darkling beetles. It is a worldwide pest of stored products, particularly food grains, and a model organism for ethological and food safety re ...


Termites

Termites are the most devastating type of book eating pest. They will eat almost every part of a book including paper, cloth, and cardboard, not to mention the damage that can be done to shelves. Termites can make entire collections unusable before the infestation is even noticed. * Powderpost termite * Western drywood termite


Ants

Some species of ants can damage books in a way that is similar to termites. * Black carpenter ant *'' Camponotus obscuripes'' * Hercules ant


Moths

Moths that feed on cloth will also feed on bookbindings, decaying organic material (which includes paper), and mold.


Fungus moths

*
Carpet moth ''Trichophaga tapetzella'', the tapestry moth or carpet moth, is a moth of the family Tineidae. It is found worldwide. The wingspan is 14–18 mm. The head is white, the forewings ochreous-white, thinly strigulated with grey; basal 2/5 dar ...
*
Case-bearing clothes moth ''Tinea pellionella'', the case-bearing clothes moth, is a species of Tineoidea, tineoid moth in the family Tineidae, the fungus moths. This species has a cosmopolitan distribution, occurring nearly worldwide.Cheema, P. S. (1956).Studies on the b ...
*
Clothing moth Clothes moth or clothing moth is the common name for several species of moth considered to be pests, whose larvae eat animal fibres (hairs), including clothing and other fabrics. These include: * ''Tineola bisselliella'', the common clothes moth ...


Pyralid moths

* Mediterranean flour moth *
Indianmeal moth The Indianmeal moth (''Plodia interpunctella''), also spelled Indian meal moth and Indian-meal moth, is a pyraloid moth of the family Pyralidae. Alternative common names are weevil moth, pantry moth, flour moth or grain moth. The almond moth ...
* Warehouse moth


Concealer moths

* brown house moth


Cockroaches

Bookdamaging cockroach species chew away at the starch in cloth bindings and paper. Their droppings can also harm books.


Wood cockroaches

*
German cockroach The German cockroach (''Blattella germanica''), colloquially known as the croton bug, is a species of small cockroach, typically about long. In color it varies from tan to almost black, and it has two dark, roughly parallel, streaks on the prono ...


Household cockroaches

*
American cockroach The american cockroach (''Periplaneta americana'') is the largest species of common cockroach, and often considered a pest. In certain regions of the U.S. it is colloquially known as the waterbug, though it is not a true waterbug since it is not ...
*
Oriental cockroach __NOTOC__ The oriental cockroach (''Blatta orientalis''), also known as the waterbug (as they live in damp areas) or black cockroaches (as their bodies are mostly dark), is a large species of cockroach, adult males being and adult females being ...
*
Smokybrown cockroach The smokybrown cockroach (''Periplaneta fuliginosa'') is a large species of cockroach, winged, and growing to a length of .Appel, A. G., & Smith II, L. M. (2002). Biology and management of the smokybrown cockroach. ''Annual Review of Entomology' ...
*
Australian cockroach The Australian cockroach (''Periplaneta australasiae'') is a common species of tropical cockroach, with a length of . It is brown overall, with the tegmina having a conspicuous lateral pale stripe or margin, and the pronotum (head shield) with a ...


Zygentoma Zygentoma are an order in the class Insecta, and consist of about 550 known species. The Zygentoma include the so-called silverfish or fishmoths, and the firebrats. A conspicuous feature of the order are the three long caudal filaments. The t ...

These insects consume portions of books that contain
polysaccharide Polysaccharides (), or polycarbohydrates, are the most abundant carbohydrates found in food. They are long chain polymeric carbohydrates composed of monosaccharide units bound together by glycosidic linkages. This carbohydrate can react with wa ...
s. Paper that is slightly ragged at the edges is usually the work of silverfish.


Lepismatidae Lepismatidae is a family of primitive wingless insects with about 190 described species. This family contains the two most familiar members of the order Zygentoma: the silverfish (''Lepisma saccharinum'') and the firebrat (''Thermobia domestica'' ...

*
Firebrat The firebrat (''Thermobia domestica'') is a small hexapod (typically 1–1.5 cm), in the order Zygentoma. Habitat Firebrats prefer higher temperatures and require some humidity, and can be found in bakeries and near boilers or f ...
*
Silverfish The silverfish (''Lepisma saccharinum'') is a species of small, primitive, wingless insect in the order Zygentoma (formerly Thysanura). Its common name derives from the insect's silvery light grey colour, combined with the fish-like appearance ...


Management

Pesticides Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and lampric ...
can be used to protect books from these insects, but they are often made with harsh chemicals that make them an unattractive option. Museums and universities that want to keep their archives bookworm free without using pesticides often turn towards temperature control. Books can be stored at low temperatures that keep eggs from hatching, or placed in a deep-freezer to kill larvae and adults. The idea was taken from commercial food storage practices, as they are often dealing with the same pests.
Pseudoscorpions Pseudoscorpions, also known as false scorpions or book scorpions, are small, scorpion-like arachnids belonging to the order Pseudoscorpiones, also known as Pseudoscorpionida or Chelonethida. Pseudoscorpions are generally beneficial to humans sin ...
such as '' Chelifer cancroides'' may live in books and feed on book-eating insects, controlling their numbers.


Idiom

The term is also used
idiom An idiom is a phrase or expression that typically presents a figurative, non-literal meaning attached to the phrase; but some phrases become figurative idioms while retaining the literal meaning of the phrase. Categorized as formulaic language, ...
atically to describe an avid or voracious reader, or a
bibliophile Bibliophilia or bibliophilism is the love of books. A bibliophile or bookworm is an individual who loves and frequently reads and/or collects books. Profile The classic bibliophile is one who loves to read, admire and collect books, often ama ...
. In its earliest iterations, it had a negative
connotation A connotation is a commonly understood cultural or emotional association that any given word or phrase carries, in addition to its explicit or literal meaning, which is its denotation. A connotation is frequently described as either positive o ...
, referring to someone who would rather read than participate in the world around them. Over the years its meaning has drifted in a more positive direction.


See also

*
Bibliomania Bibliomania can be a symptom of obsessive–compulsive disorder which involves the collecting or even hoarding of books to the point where social relations or health are damaged. Bibliomania is not to be confused with bibliophilia, which is the ...
*
Book collecting Book collecting is the collecting of books, including seeking, locating, acquiring, organizing, cataloging, displaying, storing, and maintaining whatever books are of interest to a given collector. The love of books is ''bibliophilia'', and someo ...
*
Tsundoku ''Tsundoku'' ( ja, 積ん読) refers to the phenomenon of acquiring reading materials but letting them pile up in one's home without reading them. It is also used to refer to books ready for reading later when they are on a bookshelf. The ter ...


References


Citations


Further reading


"John Francis Xavier O'Conor, Facts about bookworms: their history in literature and work in libraries (New York: Francis P. Harper, 1898.)
*Dr. John V. Richardson Jr., Ph

* ttp://www.termite.com.au/borers-pest-control.html "Timber Borers – Anobium & Lyctus Borers"br>"Study on integrated pest management for libraries and archives" – prepared by Thomas A Parker for the General Information Programme and UNISIST (Paris: Unesco, 1988)
{{Authority control Book terminology Books Insect common names Pest insects