The ruffed grouse (''Bonasa umbellus'') is a medium-sized
grouse
Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondr ...
occurring in forests from the
Appalachian Mountains
The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
across Canada to
Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
. It is the most widely distributed game bird in North America. It is
non-migratory
Bird migration is the regular seasonal movement, often north and south along a flyway, between breeding and wintering grounds. Many species of bird migrate. Migration carries high costs in predation and mortality, including from hunting by ...
. It is the only species in the genus ''Bonasa''.
The ruffed grouse is sometimes incorrectly referred to as a "
partridge
A partridge is a medium-sized Galliformes, galliform bird in any of several genera, with a wide Indigenous (ecology), native distribution throughout parts of Europe, Asia and Africa. Several species have been introduced to the Americas. They a ...
", an unrelated
phasianid
The Phasianidae are a family of heavy, ground-living birds, which includes pheasants, partridges, junglefowl, chickens, turkeys, Old World quail, and peafowl. The family includes many of the most popular gamebirds. The family is a large one and i ...
, and occasionally confused with the
grey partridge
The grey partridge (''Perdix perdix''), also known as the gray-legged partridge, English partridge, Hungarian partridge, or hun, is a gamebird in the pheasant family Phasianidae of the order Galliformes, gallinaceous birds. The scientific name ...
, a bird of open areas rather than woodlands.
[
The ruffed grouse is the state game bird of ]Pennsylvania
Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
, United States.
Taxonomy
''Bonasa umbellus'' was first described by Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1766 12th edition of ''Systema Naturae''.[ He classified it as ''Tetrao umbellus'', placing it in a subfamily with Eurasian grouse. The genus ''Bonasa'' was applied by British naturalist John Francis Stephens in 1819.][ Ruffed grouse is the preferred common name because it applies only to this species. Misleading vernacular names abound, however, and it is often called partridge (sometimes rendered pa'tridge, or shortened to pat),][ pheasant, or prairie chicken, all of which are properly applied to other birds.][ Other nicknames for ruffed grouse include drummer or thunder-chicken.][
There are 14 recognized subspecies of ruffed grouse:][
* ''B. u. yukonensis'' Grinnell, 1916
* ''B. u. umbelloides'' (]Douglas
Douglas may refer to:
People
* Douglas (given name)
* Douglas (surname)
Animals
*Douglas (parrot), macaw that starred as the parrot ''Rosalinda'' in Pippi Longstocking
*Douglas the camel, a camel in the Confederate Army in the American Civil W ...
, 1829)
* ''B. u. labradorensis'' Ouellet, 1991
* ''B. u. castanea'' Aldrich & Friedmann
Friedman, Friedmann, and Freedman are surnames of German origin, and from the 17th century were also adopted by Ashkenazi Jews. It is the 9th most common surname in Israel (8th among Jews) and most common exclusively Ashkenazi name.
They may refer ...
, 1943
* ''B. u. affinis'' Aldrich & Friedmann, 1943
* ''B. u. obscura'' Todd
Todd or Todds may refer to:
Places
;Australia:
* Todd River, an ephemeral river
;United States:
* Todd Valley, California, also known as Todd, an unincorporated community
* Todd, Missouri, a ghost town
* Todd, North Carolina, an unincorporated ...
, 1947
* ''B. u. sabini'' (Douglas, 1829)
* ''B. u. brunnescens'' Conover Conover may refer to:
People
* Conover (surname)
Places in the United States
* Conover, Iowa, a ghost town
* Conover, North Carolina, a city
* Conover, Ohio, an unincorporated community
* Conover, Wisconsin, a town
* Conover (community), Wiscon ...
, 1935
* ''B. u. togata'' (Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1766)
* ''B. u. mediana'' Todd, 1940
* ''B. u. phaios'' Aldrich & Friedmann, 1943
* ''B. u. incana'' Aldrich & Friedmann, 1943
* ''B. u. monticola'' Todd, 1940
* ''B. u. umbellus'' (Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1766)
Description
Ruffed grouse are chunky, medium-sized birds that weigh from , measure from in length and span across their short, strong wings.[ They have two distinct morphs: grey and brown. In the grey morph, the head, neck, and back are grey-brown; the breast is light with barring. There is much white on the underside and flanks. Overall, the birds have a variegated appearance; the throat is often distinctly lighter. The tail is essentially the same brownish grey, with regular barring and a broad black band near the end ("subterminal"). Brown-morph birds have tails of the same color and pattern. However, the rest of the plumage is much more brown, giving the appearance of a more uniform bird with less light plumage below and a conspicuously grey tail. There are all sorts of intergrades between the most typical morphs; warmer and more ]humid
Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present.
Humidity depen ...
conditions favor browner birds in general.
The ruffs are on the sides of the neck in both sexes. They also have a crest on top of their head, which sometimes lies flat. Both sexes are similarly marked and sized, making them difficult to tell apart, even in hand. The female often has a broken subterminal tail band. At the same time, males tend to have unbroken tail bands, though the opposite of either can occur. Females may also do a display similar to the male. Another fairly accurate sign is that rump feathers with a single white dot indicate a female; rump feathers with more than one white dot indicate a male.
The average life span of a ruffed grouse is one year, although some birds are thought to live for as long as eleven years. Ruffed grouse are polygynous
Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women.
Incidence
Polygyny is more widespread in Africa than in any ...
, and males may mate with several females during the breeding season.
Ecology
Like most grouse, they spend most of their time on the ground; mixed woodland rich in aspen
Aspen is a common name for certain tree species; some, but not all, are classified by botanists in the section ''Populus'', of the ''Populus'' genus.
Species
These species are called aspens:
*'' Populus adenopoda'' – Chinese aspen (China ...
seems to be particularly well-liked. These birds forage on the ground or in trees. They are omnivore
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
s, eating buds, leaves, berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
, seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s, and insects. According to nature writer Don L. Johnson:
Hunting
Ruffed grouse frequently seek gravel
Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone.
Gravel is classifi ...
and clover
Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus ...
along road beds during early morning and late afternoon. These are good areas to walk during this time to flush birds. Also, grouse use sandy roadbeds to dust their feathers to rid themselves of skin pests. Dusting sites are visible as areas of disturbed soils with some signs of feathers. Birds will return to these spots during the late afternoon to bathe in dust and socialize and mate.
Minnesota is the top ruffed grouse–producing state in the U.S.
Behavior
The ruffed grouse differs from other grouse species in its courtship display
A courtship display is a set of display behaviors in which an animal, usually a male, attempts to attract a mate; the mate exercises choice, so sexual selection acts on the display. These behaviors often include ritualized movement ("dances"), ...
. The ruffed grouse relies entirely on a non-vocal acoustic display, known as drumming, unlike other grouse species. The drumming itself is a rapid, wing-beating display that creates a low-frequency sound, starting slow and speeding up (thump ... thump ... thump..thump-thump-thump-thump). Even in thick woods, this can be heard for or more.
The ruffed grouse spends most of its time quietly on the ground, and when surprised, may explode into flight, beating its wings very loudly. They will burrow into the snow for warmth in the winter and may suddenly burst out of the snow when approached too closely.
The male grouse proclaims his property by engaging in a “drumming” display. This sound is made by beating his wings against the air to create a vacuum. It usually stands on a log, stone or mound of dirt when drumming. It does not strike the log to make the noise, it only uses the “drumming log” as a sort of stage.
The ruffed grouse population has a cycle, and follows the cycle no matter how much or how little hunting there is. The cycle has puzzled scientists for years, and is simply referred to as the "grouse cycle". In spite of this historical cycle, populations have been declining in Pennsylvania and management plans adopted. Habitat loss has been a concern for the species, but the introduction of the West Nile virus
West Nile virus (WNV) is a single-stranded RNA virus that causes West Nile fever. It is a member of the family ''Flaviviridae'', from the genus ''Flavivirus'', which also contains the Zika virus, dengue virus, and yellow fever virus. The virus ...
has been seen to be further increasing mortality.
See also
* State birds
* Sage grouse
Sage-grouse are grouse belonging to the bird genus ''Centrocercus.'' The genus includes two species: the Gunnison grouse (''Centrocercus minimus'') and the greater sage-grouse (''Centrocercus urophasianus''). These birds are distributed through ...
*Upland hunting Upland hunting is an American term for a form of hunting in which the hunter pursues upland birds including quail, pheasant, grouse, woodcock, prairie chicken, chukar, grey partridge, and other landfowls. Unlike aquatic birds (waterfowls and sho ...
References
Further reading
*
* Ohio Ornithological Society (2004)
Annotated Ohio state checklist
* State Symbols of Pennsylvania: State Bird, The Ruffed Grous
PDF fulltext
External links
*
*
– Cornell Lab of Ornithology
Ruffed grouse hen video Appalachian Mountains, Floyd Virginia
*
{{Authority control
Tetraonini
Symbols of Pennsylvania
Birds of the United States
Birds of Canada
Birds described in 1766
Taxa named by Carl Linnaeus
Extant Pleistocene first appearances