The bobak marmot (''Marmota bobak''), also known as the steppe marmot, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

marmot Marmots are large ground squirrels in the genus ''Marmota'', with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, w ...

that inhabits the steppes of Eastern Europe and Central Asia. It is a social animal and inhabits steppe grassland, including cultivated field borders. It hibernates for more than half the year. Litter sizes average about five offspring and it takes three years for the young marmots to reach sexual maturity. Male offspring leave the home colony after their second winter, and about 60% of mature females give birth in any one year.

Taxonomy

There are two recognized

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

: *''M. b. bobak'' — western part of range *''M. b. tschaganensis'' (alternatively ''shaganensis'') — eastern part of range Although western animals are larger and darker than eastern, their separation into subspecies is questionable, as the variation appears to be clinal. In the past, the other relatively short-furred and short-tailed marmots of the

Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...

region, i.e. Himalayan, Tarbagan,

gray Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be composed o ...

and

forest-steppe A forest steppe is a temperate-climate ecotone and habitat type composed of grassland interspersed with areas of woodland or forest. Locations Forest steppe primarily occurs in a belt of forest steppes across northern Eurasia from the eastern ...

, all were regarded as subspecies of the bobak marmot. Today they are consistently recognized as separate from the bobak marmot. Genetic studies have shown that the bobak marmot forms a

species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...

with the gray and forest-steppe marmots, while the Himalayan and Tarbagan marmots are more distantly related.

Distribution

The bobak marmot is native to the

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...

s of Eastern Europe and Central Asia, ranging from central and eastern

Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...

, through far southwestern

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

, to northern and central

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...

. Historically, its range was continuous, but it has disappeared from several regions, including its past range in southeastern

Belarus Belarus,, , ; alternatively and formerly known as Byelorussia (from Russian ). officially the Republic of Belarus,; rus, Республика Беларусь, Respublika Belarus. is a landlocked country in Eastern Europe. It is bordered by R ...

and eastern

Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...

, making its overall range discontinuous. In some places it has been reintroduced and is slowly expanding its range again. In Kazakhstan's highlands the bobak marmot comes into contact with the gray marmot and animals tend to show intermediate features. Scientists Alyona Koshkina, Johannes Kamp, et al. used satellite imagery to map the location of 7,000 Bobak marmot burrows across Kazakhstan's steppes in 2019, which are home to approximately six million marmots.

Ecology

The bobak marmot is often described as a large analog of the North American

prairie dog Prairie dogs (genus ''Cynomys'') are herbivorous burrowing ground squirrels native to the grasslands of North America. Within the genus are five species: black-tailed, white-tailed, Gunnison's, Utah, and Mexican prairie dogs. In Mexico, p ...

, with a round stomach, stubby legs, and a short tail. Bobak marmots inhabit steppe ecosystems and prosper on rolling

grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...

s and on the edge of cultivated fields.Animal Diversity Web, University of Michigan, Marmota bobak Active for about five and a half months each year, dispersers leave their natal social group after their second

hibernation Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...

. Litter sizes average a little over five, and it takes at least three years to reach sexual maturity. About 60% of adult females breed in a given year. They have a single alarm call, but studies have demonstrated that bobak marmots call faster when they live in steep terrain and slower when they live in flatter terrain. Bobak marmots' fur is used to make hats and the occasional coat. Outside

Moscow Moscow ( , US chiefly ; rus, links=no, Москва, r=Moskva, p=mɐskˈva, a=Москва.ogg) is the capital and largest city of Russia. The city stands on the Moskva River in Central Russia, with a population estimated at 13.0 million ...

, a fur-farm is experimenting with breeding bobak marmots in captivity for captive fur production. Like other marmots, the bobak is susceptible to infection by

bubonic plague Bubonic plague is one of three types of plague caused by the plague bacterium (''Yersinia pestis''). One to seven days after exposure to the bacteria, flu-like symptoms develop. These symptoms include fever, headaches, and vomiting, as well a ...

. A population of bobaks living in the

Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...

is believed to have served as a

reservoir host In Infection, infectious disease ecology and epidemiology, a natural reservoir, also known as a disease reservoir or a reservoir of infection, is the population of organisms or the specific environment in which an infectious pathogen naturally li ...

for the bubonic plague

epidemic An epidemic (from Ancient Greek, Greek ἐπί ''epi'' "upon or above" and δῆμος ''demos'' "people") is the rapid spread of disease to a large number of patients among a given population within an area in a short period of time. Epidemics ...

that struck western Russia at the end of the 19th century. Marmots are preyed upon by

snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a Felidae, felid in the genus ''Panthera'' native to the mountain ranges of Central Asia, Central and South Asia. It is listed as Vulnerable species, Vulnerable on the IUCN Red ...

s. They can also act as a buffer against other

prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...

of the snow leopard.

Cultural references

The bobak marmot is a symbol of

Luhansk Oblast Luhansk Oblast ( uk, Луга́нська о́бласть, translit=Luhanska oblast; russian: Луганская область, translit=Luganskaya oblast; also referred to as Luhanshchyna, uk, Луга́нщина) is the easternmost oblast ...

, as is found on its coat of arms and on the coats of arms of some of its districts. Bobak is on the flag and c.o.a of Bizhbulyaksky District,

Bashkortostan The Republic of Bashkortostan or Bashkortostan ( ba, Башҡортостан Республикаһы, Bashqortostan Respublikahy; russian: Республика Башкортостан, Respublika Bashkortostan),; russian: Респу́блик� ...

, Russia and on the c.o.a. of

Kupyansk Kupiansk ( uk, Куп'янськ, ) is a city in Kharkiv Oblast, Ukraine. It serves as the administrative center of Kupiansk Raion. It is also an important railroad junction for the oblast. Kupiansk hosts the administrative offices of Kupiansk Ur ...

, Ukraine

References

* Thorington, R. W. Jr. and R. S. Hoffman. 2005. Family Sciuridae. Pp. 754–818 ''in'' Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns Hopkins University Press, Baltimore. {{DEFAULTSORT:marmot, bobak

bobak marmot The bobak marmot (''Marmota bobak''), also known as the steppe marmot, is a species of marmot that inhabits the steppes of Eastern Europe and Central Asia. It is a social animal and inhabits steppe grassland, including cultivated field borders. ...

Mammals of Central Asia Rodents of Europe

Taxonomy

Distribution

Ecology

Cultural references

See also

References