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''Boa imperator'' or ''Boa constrictor imperator'' (in common usage) is a large, heavy-bodied, nonvenomous
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of snake, of the boa genus, that is commonly kept in captivity. ''Boa imperator'' is part of the family
Boidae The Boidae, commonly known as boas or boids, are a family of nonvenomous snakes primarily found in the Americas, as well as Africa, Europe, Asia, and some Pacific Islands. Boas include some of the world's largest snakes, with the green anaconda ...
and is found in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
west of the
Andes Mountains The Andes, Andes Mountains or Andean Mountains (; ) are the List of mountain ranges#Mountain ranges by length, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range i ...
(primarily
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
). It is commonly called the common boa, northern boa, Colombian boa and common northern boa and is frequently and erroneously referred to as the red-tailed boa or Colombian red-tailed boa, especially in the pet trade.


Description

''Boa imperator'' is wide-ranging, with animals living in both Central America and northern parts of South America. As a result, the appearance of this snake varies greatly depending on the specific locality. As one of the smaller ''
Boa Kwon Bo-ah (; born November 5, 1986), known professionally as BoA, is a South Korean singer, songwriter, dancer, record producer and actress. One of the most successful and influential Korean entertainers, she has been dubbed the " Queen of K- ...
'' species,Cotswold Wildlife Park and Gardens
they average between 1.3 m (4.2 ft.) and 2.5 m (8.2 ft.) in length when fully grown, but have been known to reach 3.7 m (12 ft.). They usually weigh around 6 kg (13 lb), although females are significantly larger than males. Lifespan in the wild is around 20–30 years, but 40 can be exceeded in captivity. Although ''Boa imperator'' exhibits almost identical patterns to ''
Boa constrictor The boa constrictor (scientific name also ''Boa constrictor''), also called the red-tailed boa, is a species of large, non-venomous, heavy-bodied snake that is frequently kept and bred in captivity. The boa constrictor is a member of the family B ...
'', this species often has a darker tail, usually dark brown or very dark red. They are, however, usually just as colorful as their counterparts and, like the larger boas, can be bred into a variety of different colors, given the right conditions to breed. Notably, the species is one of only two in snakes to have a confirmed XY sex chromosome system. One population is found on the
Cayos Cochinos The Cayos Cochinos or Cochinos Cays consist of two small islands (''Cayo Menor'' and ''Cayo Grande'') and 13 smaller coral cays situated northeast of La Ceiba on the northern shores of Honduras. Although geographically separate, they belong to ...
(Hog Islands) off the northern shore of
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...
. These are naturally hypomelanistic, which means that they have reduced
melanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
, thus are more lightly colored, although they retain the distinctive darker tail that is characteristic of most members of this species. The color of the tail may vary from salmon-pink to orange. Another well known population of ''Boa imperator'' is the population from
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...
. While not as small as the dwarf insular island populations, adults are still smaller than the larger ''B. c. constrictor''. The size of a mature female Nicaraguan boa is 1.1 m and 1.9 kg, while the larger female ''B. c. constrictor'' is not mature until she reaches 1.6 m and 4.5 kg. Nicaraguan individuals typically have a compact saddle pattern on their backs that is often circular in shape. These boas have a reputation for being "nippy", with some individuals being quick to bite in self-defense. Mainland specimens from
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
can be among the larger boas, but this subspecies also includes a number of dwarf insular populations, such as those from various
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
islands. These populations represent the smallest members of the species.


Scalation

''Boa imperator'' has 55–79 dorsal scales, 225–253 ventral scales, 47–69 subcaudal scales, 18–22 supralabial scales and 1–2 anal scales.


Identification

''Boa imperator'' is commonly confused with other ''Boa'' species, such as ''Boa constrictor''.


Taxonomy

''Boa imperator'' was formerly classified as a subspecies of ''Boa constrictor'' until DNA sequencing identified ''B. c. imperator'' as a separate genetic lineage with 5-7% divergence from ''B. constrictor''. Widespread acceptance is still pending, but several publications have acknowledged the new name since this finding. ''Boa imperator'' from the Pacific Coast of Mexico has been separated as another species, ''
Boa sigma ''Boa sigma'', known commonly as the Mexican west coast boa constrictor, is a species of snake in the family Boidae. The species is endemic to western Mexico. ''Boa sigma'' has previously been regarded as conspecific with ''Boa constrictor'', and ...
''. Certain ''Boa'' populations, such as Hog Island, Corn Island, Tarahumara, Ecuador, etc., are currently classified as different 'locales' of ''Boa imperator''. These locales have yet to be reclassified as official subspecies.


Geographic range

''Boa imperator'' can be found in some regions of
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and northwestern
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
, as well as several islands along the coasts of these areas. The type locality given is "l'Amerique meridionale, principalement au Mexique" (Central America, principally Mexico). ''Boa imperator'' prefers to live in
rainforests Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
due to humidity, temperature, cover from potential predators and ample prey.


Behavior

''Boa imperator'' generally live on their own, and do not interact with any other snakes unless they want to mate. They are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
, but they may bask during the day when night-time temperatures are too low. As semi-arboreal snakes, young ''Boa imperator'' may climb into trees and shrubs to forage; however, they become mostly
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
as they become older and heavier.Mehrtens JM. 1987. ''Living Snakes of the World in Color''. New York: Sterling Publishers. . ''Boa imperator''s strike when they perceive a threat. Their bite can be painful, especially from large snakes, but is rarely dangerous to humans. Specimens from Central America are more irascible, hissing loudly and striking repeatedly when disturbed, while those from South America tame down more readily.Stidworthy J. 1974. ''Snakes of the World''. Grosset & Dunlap Inc. . Like all snakes, ''Boa imperator''s in a shed cycle are more unpredictable, because the substance that lubricates between the old skin and the new makes their eyes appear milky, blue or opaque, so that the snake cannot see very well, causing it to be more defensive than it might be otherwise.


Hunting and diet

''Boa imperator'', like most ''Boa'' species, has a varied diet that consists mainly of
mammals Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s and
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s. The size of the prey item increases as it ages. ''Boa imperator'', like other ''Boa'' species, are crepuscular ambush predators. They use
constriction Constriction is a method used by various snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake i ...
as the primary means of incapacitating their prey.


Captivity

''Boa imperator'' is one of the most common snakes kept in captivity; this is mainly due to their calm dispositions, impressive size potential and variety of color and pattern choices. Captive common boas often tolerate being handled for extended periods.Reptiles Magazine Boa Care sheet
/ref> Captive ''Boa imperator'' are generally fed pre-killed rodents in an attempt to reduce damage to the specimen from the prey and for the sake of being more humane. This snake species has been a common species in the global pet trade since the 1980s, with 115,131 individuals being exported between 1989 and 2000. Global Invasive Species Database (GISD) 2015. Species profile ''Boa constrictor imperator''. Available from: http://www.iucngisd.org/gisd/species.php?sc=1439 ccessed 23 October 2018/ref> Wild caught specimens will often contain parasites, both internal and external. The most common parasite is '' Ophionyssus natricis'' or the "reptile mite". ''Boa imperator'' captive breeders will often breed for a specific color or "morph". There are several color and pattern morphs available in the pet trade, such as
albino Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and pink or blue eyes. Individuals with the condition are referred to as albino. Varied use and interpretation of the term ...
, hypomelanistic, motley and jungle individuals.


Notes


References


External links

* {{Taxonbar, from=Q58741252, from2=Q3626797
imperator The Latin word ''imperator'' derives from the stem of the verb la, imperare, label=none, meaning 'to order, to command'. It was originally employed as a title roughly equivalent to ''commander'' under the Roman Republic. Later it became a part o ...
Snakes of Central America Reptiles of Guatemala Reptiles of Mexico Reptiles of Panama Reptiles described in 1803 Taxa named by François Marie Daudin