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The blue wildebeest (''Connochaetes taurinus''), also called the common wildebeest, white-bearded gnu or brindled gnu, is a large
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mamm ...
and one of the two species of wildebeest. It is placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
'' Connochaetes'' and
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Bovidae The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species ...
, and has a close taxonomic relationship with the black wildebeest. The blue wildebeest is known to have five subspecies. This broad-shouldered antelope has a muscular, front-heavy appearance, with a distinctive, robust muzzle. Young blue wildebeest are born tawny brown, and begin to take on their adult coloration at the age of 2 months. The adults' hues range from a deep slate or bluish-gray to light gray or even grayish-brown. Both sexes possess a pair of large curved horns. The blue wildebeest is a
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
, feeding primarily on short grasses. It forms herds which move about in loose aggregations, the animals being fast runners and extremely wary. The mating season begins at the end of the rainy season and a single calf is usually born after a gestational period of about 8.5 months. The calf remains with its mother for 8 months, after which it joins a juvenile herd. Blue wildebeest are found in short-grass plains bordering bush-covered acacia savannas in southern and eastern Africa, thriving in areas that are neither too wet nor too arid. Three African populations of blue wildebeest take part in a long-distance migration, timed to coincide with the annual pattern of rainfall and grass growth on the short-grass plains where they can find the nutrient-rich forage necessary for lactation and calf growth. The blue wildebeest is native to
Angola , national_anthem = "Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label= Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kal ...
,
Eswatini Eswatini ( ; ss, eSwatini ), officially the Kingdom of Eswatini and formerly named Swaziland ( ; officially renamed in 2018), is a landlocked country in Southern Africa. It is bordered by Mozambique to its northeast and South Africa to its ...
,
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Mala ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are ...
, and
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
. Today, it is extinct in
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northe ...
, but has been successfully reintroduced in
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
. The southern limit of the blue wildebeest range is the
Orange River The Orange River (from Afrikaans/ Dutch: ''Oranjerivier'') is a river in Southern Africa. It is the longest river in South Africa. With a total length of , the Orange River Basin extends from Lesotho into South Africa and Namibia to the nort ...
, while the western limit is bounded by
Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after ...
and Mount Kenya. The blue wildebeest is widespread and is being introduced into private game farms, reserves, and conservancies. So, the International Union for Conservation of Nature and Natural Resources rates the blue wildebeest as being of least concern. The population has been estimated to be around 1.5 million, and the population trend is stable.


Taxonomy and naming

The blue wildebeest was first described by English naturalist William John Burchell in 1823 and he gave it the scientific name ''Connochaetes taurinus''. It shares the genus '' Connochaetes'' with the black wildebeest (''C. gnou''), and is placed in the family
Bovidae The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species ...
, ruminant animals with cloven hooves. The generic name ''Connochaetes'' derives from the Greek words κόννος, kónnos, "beard", and χαίτη, khaítē, "flowing hair", "mane". The specific name ''taurinus'' originates from the Greek word ''tauros'', which means a bull or bullock. The common name "blue wildebeest" refers to the conspicuous, silvery-blue sheen of the coat, while the alternative name "gnu" originates from the name for these animals used by the Khoikhoi people, a native pastoralist people of southwestern Africa. Though the blue and black wildebeest are currently classified in the same genus, the former was previously placed in a separate genus, ''Gorgon''. In a study of the
mitotic In cell biology, mitosis () is a part of the cell cycle in which replicated chromosomes are separated into two new nuclei. Cell division by mitosis gives rise to genetically identical cells in which the total number of chromosomes is mainta ...
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
s and
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondrion, mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mit ...
, which was undertaken to understand more of the evolutionary relationships between the two species, the two were found to have a close phylogenetic relationship and had diverged about a million years ago.


Subspecies

''C. taurinus'' has five subspecies: * ''C. t. taurinus'' (Burchell, 1823), the blue wildebeest, common wildebeest, or brindled gnu is found in southern Africa. Its range extends from Namibia and South Africa to Mozambique (north of the
Orange River The Orange River (from Afrikaans/ Dutch: ''Oranjerivier'') is a river in Southern Africa. It is the longest river in South Africa. With a total length of , the Orange River Basin extends from Lesotho into South Africa and Namibia to the nort ...
) and from southwestern Zambia (south of the
Zambezi River The Zambezi River (also spelled Zambeze and Zambesi) is the fourth-longest river in Africa, the longest east-flowing river in Africa and the largest flowing into the Indian Ocean from Africa. Its drainage basin covers , slightly less than hal ...
) to southern Angola. * ''C. t. johnstoni'' (Sclater, 1896), the Nyassaland wildebeest, occurs from Mozambique (north of the Zambezi River) to east-central Tanzania. It is now extinct in Malawi. * ''C. t. albojubatus'' (Thomas, 1912), the eastern white-bearded wildebeest, is found in the Gregory Rift Valley (south of the equator). Its range extends from northern Tanzania to central Kenya. * ''C. t. mearnsi'' (Heller, 1913), the western white-bearded wildebeest, is found in northern Tanzania and southern Kenya. Its range extends from the west of the Gregory Rift Valley to Speke Bay on
Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after ...
. * ''C. t. cooksoni'' (Blaine, 1914), Cookson's wildebeest, is restricted to the Luangwa Valley in Zambia. Sometimes, it may wander into the plateau region of central Malawi. In addition, the distinctive appearance of a western form, ranging from the Kalahari to central Zambia, suggests that subspecies ''mattosi'' (Blaine, 1825) may also prove distinct from subspecies ''taurinus''. The western form can be recognised even at a distance by its upright mane, long beard, and minimal brindling.


Hybrids

The blue wildebeest is known to hybridise with the black wildebeest. The differences in social behaviour and habitats have historically prevented interspecific hybridisation, but it may occur when both species are confined within the same area, and the offspring are usually fertile. A study of these hybrid animals at
Spioenkop Dam Nature Reserve Spioenkop Dam Nature Reserve or Spion Kop Nature Reserve is a protected area in KwaZulu-Natal, South Africa. It lies close to Ladysmith with Winterton being the closest town, and is about . The historic battlefield site can be reached by roa ...
in South Africa revealed that many had congenital abnormalities relating to their teeth, horns, and the Wormian bones of the skull. Another study reported an increase in the size of the hybrid as compared to either of its parents. In some hybrid animals, the auditory bullae are highly deformed, and in others, the
radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...
and
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
are fused.


Genetics and evolution

The
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respecti ...
number of
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
s in the blue wildebeest is 58. Chromosomes were studied in a male and a female wildebeest. In the female, all except a pair of very large submetacentric chromosomes were found to be acrocentric. Metaphases were studied in the male's chromosomes, and very large submetacentric chromosomes were found there as well, similar to those in the female both in size and morphology. The rest were acrocentric. The
X chromosome The X chromosome is one of the two sex-determining chromosomes ( allosomes) in many organisms, including mammals (the other is the Y chromosome), and is found in both males and females. It is a part of the XY sex-determination system and XO se ...
is a large acrocentric, while the
Y chromosome The Y chromosome is one of two sex chromosomes ( allosomes) in therian mammals, including humans, and many other animals. The other is the X chromosome. Y is normally the sex-determining chromosome in many species, since it is the presence or a ...
is a minute one. This species of wildebeest seems to have evolved around 2.5 million years ago. The black wildebeest is believed to have diverged from the blue wildebeest to become a distinct species around 1 million years ago, in the Middle to Late
Pleistocene The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed ...
. Fossil evidence suggests that blue wildebeest were quite common in the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site and is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest concentration of ...
in the past. Apart from eastern Africa, fossils are commonly found in
Elandsfontein Elandsfontein may refer to: * Elandsfontein, an archaeological site near Hopefield, South Africa Hopefield is a settlement in West Coast District Municipality in the Western Cape province of South Africa on the R45 between Malmesbury and Vredenbur ...
, Cornelia, and Florisbad.


Description

The blue wildebeest exhibits
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
, with males being larger and darker than females. The blue wildebeest is typically in head-and-body length. The average height of the species is . Males typically weigh and females weigh . A characteristic feature is the long, black tail, which is around in length. All features and markings of this species are bilaterally symmetrical for both sexes. The average life span is 20 years in the wild, and 21 years in captivity. The oldest known captive individual lived for 24.3 years.


Colouration

This broad-shouldered antelope has a muscular, front-heavy appearance, with a distinctive robust muzzle. Young are born tawny brown, and begin to take on their adult coloration at the age of 2 months. The adults' hues range from a deep slate or bluish-gray to light gray or even grayish-brown. The back and flanks are slightly lighter than the ventral surface and underparts. Dark brown, vertical stripes mark the area between the neck and the back of the ribcage, thus giving it the name "brindled gnu". The manes of both sexes appear long, stiff, thick, and jet black, the same colour as the tail and face. While the manes of the western and eastern white-bearded wildebeest are lank, those of the Nyassaland wildebeest and common wildebeest stick up. Scent glands, which secrete a clear oil, are present in the forefeet and are larger in males than females. In terms of
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
length, the smallest subspecies of the blue wildebeest is the western white-bearded wildebeest. It is also the darkest subspecies; the eastern white-bearded wildebeest is the lightest race. Both subspecies possess a creamy white beard, whereas the beard is black in both the Nyassaland wildebeest and the common wildebeest. The longest muzzles are found in the Nyassaland wildebeest, and the shortest in female western white-bearded wildebeest.


Horns

Both sexes possess a pair of large horns, which are shaped like
parentheses A bracket is either of two tall fore- or back-facing punctuation marks commonly used to isolate a segment of text or data from its surroundings. Typically deployed in symmetric pairs, an individual bracket may be identified as a 'left' or 'r ...
. These extend outward to the side, and then curve upward and inward. In the males, the horns can be long, while the horns of the females are long. Despite being an antelope, the blue wildebeest possesses various
bovine Bovines (subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship be ...
characteristics. For instance, the horns resemble those of the female African buffalo. Further, the heavy build and disproportionately large forequarters give it a bovine appearance.


Ecology and behavior

The blue wildebeest is mostly active during the morning and the late afternoon, with the hottest hours of the day being spent in rest. These extremely agile and wary animals can run at speeds up to , waving their tails and tossing their heads. An analysis of the activity of blue wildebeest at the Serengeti National Park showed that the animals devoted over half of their total time to rest, 33% to grazing, 12% to moving about (mostly walking), and a little to social interactions. However, variations existed among different age and sex groups. The wildebeest usually rest close to others of their kind and move about in loose aggregations. Males form bachelor herds, and these can be distinguished from juvenile groups by the lower amount of activity and the spacing between the animals. Around 90% of the male calves join the bachelor herds before the next mating season. Bulls become territorial at the age of four or five years, and become very noisy (most notably in the western white-bearded wildebeest) and active. The bulls tolerate being close to each other and of plain can accommodate 270 bulls. Most territories are of a temporary nature and fewer than half of the male population hold permanent territories. In general, blue wildebeest rest in groups of a few to thousands at night, with a minimum distance of between individuals (though mothers and calves may remain in contact). They are a major prey item for
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphi ...
s,
cheetah The cheetah (''Acinonyx jubatus'') is a large cat native to Africa and central Iran. It is the fastest land animal, estimated to be capable of running at with the fastest reliably recorded speeds being , and as such has evolved specialized ...
s,
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia ...
s,
African wild dog The African wild dog (''Lycaon pictus''), also called the painted dog or Cape hunting dog, is a wild canine which is a native species to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Ly ...
s,
hyena Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the clas ...
s, and crocodiles. Female calves will stay with their mothers and other related females of the herd throughout their lives. Female individuals in a herd are from a wide range of ages, from yearlings to the oldest cow. During the wet season, the females generally lead the herd towards nutritious areas of grasses and areas where predators can be avoided. This is to ensure that newborn calves have the highest chance of survival as well as gaining the most nutritious milk. Bulls mark the boundaries of their territories with heaps of dung, secretions from their scent glands, and certain behaviors. Body language used by a territorial male includes standing with an erect posture, profuse ground pawing, and horning, frequent defecation, rolling and bellowing, and the sound "ga-noo" being produced. When competing over territory, males grunt loudly, paw the ground, make thrusting motion with their horns, and perform other displays of aggression.


Diet

The blue wildebeest is a
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
, feeding primarily on the short grasses which commonly grow on light, and alkaline soils that are found in savanna grasslands and on plains. The animal's broad mouth is adapted for eating large quantities of short grass and it feeds both during the day and night. When grass is scarce, it will also eat the foliage of shrubs and trees. Wildebeest commonly associate with
plains zebra The plains zebra (''Equus quagga'', formerly ''Equus burchellii''), also known as the common zebra, is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south o ...
s as the latter eat the upper, less nutritious grass canopy, exposing the lower, greener material which the wildebeest prefer. Whenever possible, the wildebeest likes to drink twice daily and due to its regular requirement for water, it usually inhabits moist grasslands and areas with available water sources. The blue wildebeest drinks 9 to 12 litres of water every one to two days. Despite this, it can also survive in the arid
Kalahari desert The Kalahari Desert is a large semi-arid sandy savanna in Southern Africa extending for , covering much of Botswana, and parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African Namib coast ...
, where it obtains sufficient water from melons and water-storing roots and tubers. In a study of the dietary habits of the wildebeest, the animals were found to be feeding on the three dominant kinds of grass of the area, namely: '' Themeda triandra'', '' Digitaria macroblephara'', and '' Pennisetum mezianum''. The time spent grazing increased by about 100% during the dry season. Though the choice of the diet remained the same in both the dry and the wet season, the animals were more selective during the latter.


Reproduction

Male blue wildebeest become sexually mature at about 2 years of age, while females can conceive at 16 months if adequately nourished. Nevertheless, most females do not start to breed until a year later. The mating season, which lasts for about 3 weeks, coincides with the end of the rainy season. This means that the animals are in good condition, having been feeding on highly nutritious new grass growth, and the conception rate is often as high as 95%. The mating season, or rut, typically begins on the night of a full moon, suggesting that the lunar cycle influences breeding. At this time,
testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteris ...
production peaks in males, resulting in increased calling and territorial behavior. The activities of these sexually excited males may also stimulate the female to come into estrus. As they stake out their territories and compete for females, males exhibit rivalry. When they clash, they face up to each other with bent knees and exchange horn thrusts. Elaborate individual displays are made during their rivalry, and they may bellow, snort, and dig their horns into the ground. Once dominance has been established, each male attempts to lure the female into his domain. During
courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private ...
, urination and low-stretch are common activities, and the male soon attempts to mount the female. A receptive female holds her tail to one side and stands still while copulation takes place. Matings may be repeated several times and may take place twice or more times within a minute. The male neither eats nor rests when a female is present in his territory, and during this time, the female keeps close to the male, often rubbing her head on his torso and sniffing his penis. While in season, a female may visit several territories and mate with several different males. The gestation period is about 8.5 months, and between 80 and 90% of the calves are born within a 3-week time period. Female wildebeest give birth in the middle of a herd rather than alone, and typically in the middle of the day. This allows time for the newborn to become steady on its feet before night falls and the predators become more active. Calves weigh about at birth, and can usually stand on their own within a few minutes of birth. To escape predation, calves remain close to their mothers for a significant time, and may continue suckling until the next year's calf is nearly due. Male calves leave their mother at about 8 months and form herds with other male juveniles. In large female herds, 80% of the wildebeest offspring survive the first month, compared to a 50% survival rate in smaller herds.


Diseases and parasites

The blue wildebeest is susceptible to
foot-and-mouth disease Foot-and-mouth disease (FMD) or hoof-and-mouth disease (HMD) is an infectious disease, infectious and sometimes fatal virus (biology), viral disease that affects cloven-hoofed animals, including domestic and wild Bovidae, bovids. The virus causes ...
,
anthrax Anthrax is an infection caused by the bacterium '' Bacillus anthracis''. It can occur in four forms: skin, lungs, intestinal, and injection. Symptom onset occurs between one day and more than two months after the infection is contracted. The s ...
, sarcoptic mange, and hoof gangrene. The herpesvirus was first isolated from the blue wildebeest in 1960 by veterinary scientist Walter Plowright. Although the causes of death vary from year to year, in one drought in Botswana, young calves and aged females were the most likely to die. On another occasion, an estimated 47% of deaths were caused by disease, 37% were due to predation, and the remainder were the result of accidents. The animal can be host to a number of different parasites. In one study, blue wildebeest were found to be hosts to 13 species of nematodes, one
trematode Trematoda is a Class (biology), class of flatworms known as flukes. They are obligate parasite, obligate internal Parasitism, parasites with a complex biological life cycle, life cycle requiring at least two Host_(biology), hosts. The intermedia ...
, larvae of five oestrid flies, three species of
lice Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been recognized as an order, infraorder, or a parvorder, as a resul ...
, seven ixodid tick species, one
mite Mites are small arachnids (eight-legged arthropods). Mites span two large orders of arachnids, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari, but genetic analysis does not show clear evid ...
, and the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
e of a
tongue worm The Pentastomida are an enigmatic group of parasitic arthropods commonly known as tongue worms due to the resemblance of the species of the genus ''Linguatula'' to a vertebrate tongue; molecular studies point to them being degenerate crustaceans ...
. Of these, most were more prevalent at some times of the year than others. Generally, the larvae of '' Gedoelstica'' and ''Oestrus'' occur in the
nasal passage The nasal cavity is a large, air-filled space above and behind the nose in the middle of the face. The nasal septum divides the cavity into two cavities, also known as fossae. Each cavity is the continuation of one of the two nostrils. The nasal c ...
s and respiratory cavities of the blue wildebeest, and sometimes migrate to the brain. Compared to some other bovids, blue wildebeest are resistant to infestations by several species of ticks.


Distribution and habitat

The blue wildebeest is native to
Kenya ) , national_anthem = " Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label= Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kal ...
,
Zambia Zambia (), officially the Republic of Zambia, is a landlocked country at the crossroads of Central, Southern and East Africa, although it is typically referred to as being in Southern Africa at its most central point. Its neighbours are ...
,
Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and Mozam ...
,
Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Mala ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring count ...
,
Eswatini Eswatini ( ; ss, eSwatini ), officially the Kingdom of Eswatini and formerly named Swaziland ( ; officially renamed in 2018), is a landlocked country in Southern Africa. It is bordered by Mozambique to its northeast and South Africa to its ...
, and
Angola , national_anthem = "Angola Avante"() , image_map = , map_caption = , capital = Luanda , religion = , religion_year = 2020 , religion_ref = , coordina ...
. Today, it is extinct in
Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northe ...
, but has been successfully reintroduced into
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and e ...
. Blue wildebeest are mainly found in short-grass plains bordering bush-covered acacia savannas in southern and eastern Africa, thriving in areas that are neither too wet nor too arid. They can be found in habitats that vary from overgrazed areas with dense bush to open
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
floodplains. Trees such as ''
Brachystegia ''Brachystegia'' is a genus of tree of the subfamily Detarioideae that is native to tropical Africa. Trees of the genus are commonly known as miombo, and are dominant in the miombo woodlands of central and southern tropical Africa. The Zambezian ...
'' and '' Combretum'' spp. are common in these areas. Blue wildebeest can tolerate arid regions as long as a potable water supply is available, normally within about distance. The southern limit of the blue wildebeest stops at the
Orange River The Orange River (from Afrikaans/ Dutch: ''Oranjerivier'') is a river in Southern Africa. It is the longest river in South Africa. With a total length of , the Orange River Basin extends from Lesotho into South Africa and Namibia to the nort ...
, while the western limit is bounded by
Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after ...
and Mount Kenya. The range does not include montane or temperate grasslands. These wildebeest are rarely found at altitudes over . With the exception of a small population of Cookson's wildebeest that occurs in the Luangwa Valley (Zambia), the wildebeest is absent in the wetter parts of the southern savanna country, and particularly is not present in miombo woodlands. Three African populations of blue wildebeest take part in long-distance migrations, timed to coincide with the annual pattern of rainfall and grass growth on the short-grass plains, where they can find the nutrient-rich forage necessary for lactation and calf growth. The timing of the migration in both directions can vary considerably from year to year. At the end of the rainy season, they migrate to dry-season areas in response to a lack of drinking water. When the rainy season begins again a few months later, the animals trek back to their wet-season range. These movements and access to nutrient-rich forage for reproduction allow migratory wildebeest populations to grow to much larger numbers than resident populations. Many long-distance migratory populations of wildebeest existed 100 years ago, but currently, all but three migrations ( Serengeti, Tarangire, and Kafue) have been disrupted, cut off, and lost.


Threats and conservation

Major human-related factors affecting populations include large-scale deforestation, the drying up of water sources, the expansion of settlements and poaching. Diseases of domestic cattle such as sleeping sickness can be transmitted to the animals and take their toll. The erection of fences that interrupt traditional migratory routes between wet and dry-season ranges have resulted in mass death events when the animals become cut off from water sources and the areas of better grazing they are seeking during droughts. A study of the factors influencing wildebeest populations in the Maasai Mara ecosystem revealed that the populations had undergone a drastic decline of around 80% from about 119,000 individuals in 1977 to around 22,000 twenty years later. The major cause of this was thought to be the expansion of agriculture, which led to the loss of wet-season grazing and the traditional calving and breeding ranges. Similarly, drastic declines have recently occurred in the Tarangire wildebeest migration. The total number of blue wildebeest is estimated to be around 1,550,000. The population trend overall is stable and the numbers in the Serengeti National Park (Tanzania) have increased to about 1,300,000. The population density ranges from 0.15/km2 in Hwange and Etosha National Parks to 35/km2 in
Ngorongoro Crater The Ngorongoro Conservation Area (, ) is a protected area and a UNESCO World Heritage Site located in Ngorongoro District, west of Arusha City in Arusha Region, within the Crater Highlands geological area of northern Tanzania. The area is n ...
and Serengeti National Park, where they are most plentiful. Blue wildebeest have also been introduced into a number of private game farms, reserves, and conservancy areas. For these reasons, the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natur ...
rates the blue wildebeest as being of least concern. However, the numbers of the eastern white-bearded wildebeest (''C. t. albojubatus'') have seen a steep decline to a current level of probably 6,000 to 8,000 animals, and this is causing some concern.


Relationship with humans

As one of the major herbivores of southern and eastern Africa, the blue wildebeest is one of the animals that draw tourists to the area to observe big game, and as such, it is of major economic importance to the region. Traditionally, blue wildebeest have been hunted for their hides and meat, the skin making good-quality leather, though the flesh is coarse, dry, and rather tough. However, blue wildebeest can also affect human beings negatively. They can compete with domestic livestock for grazing and water and can transmit fatal diseases like rinderpest to cattle and cause epidemics among animals. They can also spread ticks, lungworms,
tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass is Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Cestod ...
s, flies, and paramphistome flukes. An ancient carved slab of
slate Slate is a fine-grained, foliated, homogeneous metamorphic rock derived from an original shale-type sedimentary rock composed of clay or volcanic ash through low-grade regional metamorphism. It is the finest grained foliated metamorphic ro ...
depicting an animal very similar to the blue wildebeest has been discovered. Dating back to around 3000 BC, it was found in Hierakonopolis (Nekhen), which used to be the religious and political capital of
Upper Egypt Upper Egypt ( ar, صعيد مصر ', shortened to , , locally: ; ) is the southern portion of Egypt and is composed of the lands on both sides of the Nile that extend wikt:downriver, upriver from Lower Egypt in the north to Nubia in the south. ...
at that time. This may be evidence that the animal used to occur in North Africa and was associated with the ancient Egyptians.


References


External links


Blue Wildebeest Photo and Fact Sheet''Connochaetes taurinus''
Mammal Species of the World {{Use dmy dates, date=April 2017 blue wildebeest Fauna of East Africa Mammals of Southern Africa Extant Pleistocene first appearances blue wildebeest blue wildebeest Bovids of Africa