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''Bouteloua gracilis'', the blue grama, is a long-lived, warm-season ( C4)
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in law ...
, native to North America. It is most commonly found from
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
, Canada, east to
Manitoba , image_map = Manitoba in Canada 2.svg , map_alt = Map showing Manitoba's location in the centre of Southern Canada , Label_map = yes , coordinates = , capital = Win ...
and south across the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
, Great Plains, and U.S.
Midwest The Midwestern United States, also referred to as the Midwest or the American Midwest, is one of four census regions of the United States Census Bureau (also known as "Region 2"). It occupies the northern central part of the United States. ...
states, onto the northern
Mexican Plateau The Central Mexican Plateau, also known as the Mexican Altiplano ( es, Altiplanicie Mexicana), is a large arid-to-semiarid plateau that occupies much of northern and central Mexico. Averaging above sea level, it extends from the United States b ...
in
Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
. Blue grama accounts for most of the net primary productivity in the shortgrass prairie of the central and southern Great Plains. It is a green or greyish, low-growing, drought-tolerant grass with limited maintenance.


Description

Blue grama has green to greyish leaves less than wide and long. The overall height of the plant is at maturity. The flowering stems ( culms) are long. At the top are one to four, usually two, comb-like spikes, which extend out at a sharp angle from the flowering stem. Each spike has 20 to 90 spikelets. Each spikelet is long, and has one fertile
floret This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary ...
and one or two reduced sterile ones. Below the florets are two glumes, one long and the other long. The fertile floret has a
lemma Lemma may refer to: Language and linguistics * Lemma (morphology), the canonical, dictionary or citation form of a word * Lemma (psycholinguistics), a mental abstraction of a word about to be uttered Science and mathematics * Lemma (botany), a ...
( bract) long, with three short awns (bristles) at the tip, and the sterile floret has a lemma about long with three awns about long. If pollinated, the fertile floret produces an oblong-elliptic brown seed long. When the seed is mature, the whole spikelet detaches, except for the two glumes, which remain. The
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s generally grow outwards, and deep. Blue grama is readily established from
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
, but depends more on
vegetative reproduction Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spe ...
via
tiller A tiller or till is a lever used to steer a vehicle. The mechanism is primarily used in watercraft, where it is attached to an outboard motor, rudder post or stock to provide leverage in the form of torque for the helmsman to turn the rudder ...
s. Seed production is slow, and depends on soil moisture and temperature. Seeds dispersed by
wind Wind is the natural movement of air or other gases relative to a planet's surface. Winds occur on a range of scales, from thunderstorm flows lasting tens of minutes, to local breezes generated by heating of land surfaces and lasting a few ...
only reach a few meters (6 ft); farther distances are reached with insects, birds, and mammals as dispersal agents. Seedling establishment, survival, and growth are greatest when isolated from neighboring adult plants, which effectively exploit water in the seedling's root zone. Successful establishment requires a modest amount of soil moisture during the extension and development of adventitious roots. Established plants are
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and other ...
-, cold-, and
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
-tolerant, though prolonged drought leads to a reduction in root number and extent. They employ an opportunistic water-use strategy, rapidly using water when available, and becoming
dormant Dormant, "sleeping", may refer to: Science *Dormancy Dormancy is a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps ...
during less-favorable conditions. In terms of successional status, blue grama is a late seral to climax species. Recovery following disturbance is slow and depends on the type and extent of the disturbance.


Distribution

Blue grama has the widest distribution of all grama grasses. It will grow on most soil types, and readily adapts to local conditions. It can be found as far north as
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
, and as far south as Mexico. It is present in most of the Midwestern United States, extending east to Missouri and Texas, and as far west as Southern California. It has been introduced to some eastern states, as well as South America.


Horticulture and agriculture

Blue grama is valued as
forage Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also us ...
, and is the most valuable grama. It is an ideal range grass in the southwest. It will survive heavy grazing and extreme drought, and is quite palatable to livestock. Grazing of blue grama rangelands might also prevent invasion of undesirable
weed A weed is a plant considered undesirable in a particular situation, "a plant in the wrong place", or a plant growing where it is not wanted.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. ...
y plants. Blue grama is grown by the horticulture industry, and used in
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
gardens, naturalistic and
native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equ ...
landscaping Landscaping refers to any activity that modifies the visible features of an area of land, including the following: # Living elements, such as flora or fauna; or what is commonly called gardening, the art and craft of growing plants with a goal ...
,
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
restoration projects, and residential, civic, and highway erosion control. Blue grama flowers are also used in dried flower arrangements. Blue grama is the state grass of
Colorado Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the wes ...
and
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe, New Mexico, Santa Fe , LargestCity = Albuquerque, New Mexico, Albuquerque , LargestMetro = Albuquerque metropolitan area, Tiguex , Offi ...
. It is listed as an endangered species in
Illinois Illinois ( ) is a state in the Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolitan areas include, Peoria and Roc ...
. Blue grama has a large
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding ...
relative to other grama grasses. It can be
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respecti ...
or tetraploid. Among the Zuni people, the grass bunches are tied together and the severed end is used as a hairbrush, the other as a broom. Bunches are also used to strain goat's milk. The
Costanoan The Ohlone, formerly known as Costanoans (from Spanish meaning 'coast dweller'), are a Native American people of the Northern California coast. When Spanish explorers and missionaries arrived in the late 18th century, the Ohlone inhabited the ...
, or
Ohlone The Ohlone, formerly known as Costanoans (from Spanish meaning 'coast dweller'), are a Native American people of the Northern California coast. When Spanish explorers and missionaries arrived in the late 18th century, the Ohlone inhabited the ...
, use(d) the hollow stems as drinking straws. The
Navajo The Navajo (; British English: Navaho; nv, Diné or ') are a Native Americans in the United States, Native American people of the Southwestern United States. With more than 399,494 enrolled tribal members , the Navajo Nation is the largest fe ...
use(d) it as sheep and horse feed.


Ecology

Blue grama is the dominant species of the US shortgrass steppe ecoregion. Populations of blue grama across the great plains are genetically differentiated (even at small spatial scales) and show functional trait variance connected to climate. Blue grama from more arid grasslands are also characterized by greater phenotypic plasticity. It is a larval host to the
Garita skipperling ''Oarisma garita'', the Garita skipperling, western skipperling or Garita skipper, is a North American butterfly in the family Hesperiidae (skippers), subfamily Hesperiinae (grass skippers). This skipper ranges southeastern Manitoba to British C ...
, green skipper,
Pahaska skipper ''Hesperia pahaska'', the Pahaska skipper, is a butterfly of the family Hesperiidae. It is found from North America in a narrow belt of states extending from Texas to North Dakota, just reaching Canada in southern Saskatchewan and Manitoba.
, Rhesus skipper,
Simius roadside skipper ''Amblyscirtes simius'', the simius roadside skipper, is a butterfly of the family Hesperiidae. It is found in North America from Texas to North Dakota but has been recorded in southern Saskatchewan. The wingspan is 21–23 mm. Adults are o ...
, and the Uncas skipper.The Xerces Society (2016), ''Gardening for Butterflies: How You Can Attract and Protect Beautiful, Beneficial Insects'', Timber Press.


References


External links

{{Taxonbar, from=Q2922605 gracilis Grasses of North America Warm-season grasses of North America Native grasses of the Great Plains region Grasses of Canada Grasses of Mexico Grasses of the United States Native grasses of California Native grasses of Nebraska Native grasses of Oklahoma Native grasses of Texas Flora of Mexico Flora of the Western United States Flora of the United States Flora of the South-Central United States Flora of the California desert regions Flora of the Canadian Prairies Flora of the Chihuahuan Desert Flora of the Great Basin Flora of the Great Lakes region (North America) Flora of the Mexican Plateau Natural history of the Mojave Desert Plants described in 1816 Symbols of Colorado Symbols of New Mexico Garden plants of North America Drought-tolerant plants Flora without expected TNC conservation status