Blue-throated Hillstar
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The blue-throated hillstar (''Oreotrochilus cyanolaemus'') is a
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
found only in a small portion of the southwestern
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
in
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
. It was discovered in 2017.


Taxonomy and evolution

The species name "cyanolaemus" is based on the Latinized Greek words for ultramarine blue (kuanos) and throat (laimos). Based on a sequences of a single
mitochondrial gene Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
, ND2, the species was found to be closely related to a clade composed of the
black-breasted hillstar The black-breasted hillstar (''Oreotrochilus melanogaster'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is endemic to Peru.HBW and BirdLife International (2020) ''Handbook of the Birds of the Worl ...
(''O. melanogaster'') and the
green-headed hillstar The green-headed hillstar (''Oreotrochilus stolzmanni'') is a species of hummingbird found in the Andes of southern Ecuador and northern and central Peru.Handbook of the Birds of the World. Vol. 5, J. del Hoyo, A. Elliott & J. Sargatal (eds.), Bar ...
(''O. stolzmanni'').


Description

The top of the head of the male blue-throated hillstar is iridescent emerald green, which extends from the forehead to the base of the crown. The iridescent highlights are blue-green. The nape of the neck, mantle, lower portion of the back, rump, and upperwing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are sm ...
are bright emerald green. Some feathers in these regions have blue-green edging, and some have bronze highlights. The uppertail coverts are emerald blue-green. The upper surface of the central
flight feather Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
s are blackish blue. The other flight feathers and the upperwing coverts are dull black to dusky greenish blue, with a blue sheen. The outermost primary feathers have a white edge. The feathers of the throat and chin are a glittering
ultramarine Ultramarine is a deep blue color pigment which was originally made by grinding lapis lazuli into a powder. The name comes from the Latin ''ultramarinus'', literally 'beyond the sea', because the pigment was imported into Europe from mines in Afgh ...
blue, with white bases separated from the blue parts of the feather by a black line. The feathers of the throat have slightly green tips, and are longer on the sides of the throat than in the middle; the extent of these green tips varies between individuals. The bright blue throat patch was the basis for its name. The bird has a velvety black collar around its throat. The belly and the breast are a dull white color, and the feathers in these regions have black bases. There is a black longitudinal stripe in the middle of the belly. The sides and flanks of the bird are between green and greenish gray, with metallic green highlights. The undertail coverts are dull grayish-buff, while the lower surface of the tail is dull white. The flight feathers of the tail are blackish blue. In the female of the species, the back is emerald green, with some variation between bluish and bronze-green. The uppertail coverts are brighter and more blue than in the male. The wing coverts are dusky bluish green, and the outer primaries have a white edge. The throat and chin are speckled; the feathers are black at the base, whitish in the middle, and grayish olive on the outer half, with a dark green spot. The sides of the throat are whiter than in the male. The extent of the dusky color in the throat varies between individuals. The flanks, belly, breast, and undertail coverts are grayish buff with occasional brighter olive green feathers, as are the ventral regions of the tail. The tail has white patches, the size and shape of which vary between individuals. The flight feathers of the tail are dusky bluish green. In both sexes, the beak is black, with yellowish
tomia The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
. A
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen weighed 8.15 g. Males of the species may be distinguished from congeners by the presence of the ultramarine throat, emerald green head and upperparts with bluish-green feather tips, and emerald green tips to the throat feathers. Females may be distinguished by the presence of a dusky-gray chin and upper throat in contrast to a whitish lower throat, and emerald green head and upperparts with bluish green feather tips on the head and a bluish green shine on the body. The males are most similar to those of the
green-headed hillstar The green-headed hillstar (''Oreotrochilus stolzmanni'') is a species of hummingbird found in the Andes of southern Ecuador and northern and central Peru.Handbook of the Birds of the World. Vol. 5, J. del Hoyo, A. Elliott & J. Sargatal (eds.), Bar ...
(''O. stolzmanni'') and the
Ecuadorian hillstar The Ecuadorian hillstar or Chimborazo hillstar (''Oreotrochilus chimborazo'') is a species of hummingbird. It is native to the Andes of Ecuador and extreme southern Colombia. Its main habitat type is high-altitude mountain grassland between 3500 ...
(''O. chimborazo''); they are differentiated most clearly by the green throat patch in the green-headed hillstar, and the purple hood in the Ecuadorian hillstar. The vocalizations of the blue-throated hillstar are similar to those of its congeners; they are highly variable, and include rapid titters of rising and falling frequency, and single noted calls consisting of an "up–down stroke".


Habitat and ecology

The preferred habitat of the blue-throated hillstar is grassland with patches of shrubs in the Andean '' paramo''. In the region that the type specimen was collected, the shrubs were dominated by ''
Chuquiraga ''Chuquiraga'' is a genus of flowering plants in the family Asteraceae.''Chuquirag ...
''. The terrain in which it was found included hilly regions, steep terrain, rocky outcrops, and stream banks. ''O. cyanolaemus'' was observed to feed by clinging to branches sideways or upside down, or by perching next to flowers. It was observed feeding most frequently from ''
Chuquiraga jussieui ''Chuquiraga jussieui'', the flower of the Andes, is a species of flowering plant in the family Asteraceae. It is a low shrub, reaching a height of about 75 cm. ''Chuquiraga jussieui'' is an endangered flower. Flowers are pale yellow or oran ...
'', and also from ''Macleania rupestris'' and ''Lleresia hypoleuca''. Collected males were also found to have eaten flies. Some females were once observed making aerial sallies after swarming flies. Blue-throated hillstars were frequently observed being chased away from perches by Shining Sunbeams (''Aglaeactis cupripenni''). Aggressive interactions were also seen between it and the Viridian metaltail (''Metallura williami'').


Distribution and conservation

The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to an area of about where it is found along bush-lined creeks on small ridges not surpassing 3700 m. The range of this species is restricted by the Jubones- León river valley to the north and northeast and the Puyango- Catamayo drainage basin to the south. No more than 750 individuals have been estimated to exist, and this population is highly threatened by fire, grazing, and gold mining. Nature and Culture International is working with Ecuador’s Ministry of Environment, Ministry of Tourism, Water Secretary, municipalities of
Saraguro Saraguro (also Sarakuru) is the capital of Saraguro Canton in Loja Province, Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which litera ...
and
Zaruma Zaruma, officially Villa Real de San Antonio del Cerro de Oro de Zaruma is a town in the south of Ecuador, El Oro Province. It is located in the south-east of this province, at an altitude of 1200 metres above sea level, on an inter-Andes route w ...
, Kawsay Foundation and local and indigenous communities to establish a new conservation area for this species. The proposed area – spanning around 175,000 acres in
Cerro de Arcos The Cerro de Arcos (English: Hill of arches) is a windswept rock formation in the high páramo of the Ecuadorian southern sierra. It is situated on the border between the provinces El Oro and Loja, on the elevated plain between the Cordillera d ...
– will protect the blue-throated hillstar’s entire habitat in addition to critical
páramo Páramo () can refer to a variety of alpine tundra ecosystems located in the Andes Mountain Range, South America. Some ecologists describe the páramo broadly as "all high, tropical, montane vegetation above the continuous timberline". A narrower ...
and cloud forest ecosystems, wetlands and important water sources. The area will also preserve portions of the
Inca trail The Inca Trail to Machu Picchu (also known as ''Camino Inca'' or ''Camino Inka'') is a hiking trail in Peru that terminates at Machu Picchu. It consists of three overlapping trails: ''Mollepata'', ''Classic'', and ''One Day''. ''Mollepata'' is ...
, an ancient trail used by the
Inca empire The Inca Empire (also known as the Incan Empire and the Inka Empire), called ''Tawantinsuyu'' by its subjects, (Quechua for the "Realm of the Four Parts",  "four parts together" ) was the largest empire in pre-Columbian America. The admin ...
to connect Cuzco with its northern capital, Quito.


References

{{Taxonbar, from=Q56807945 Hillstars Endemic birds of Ecuador Birds of the Ecuadorian Andes Birds described in 2018 Taxa named by Niels Krabbe