The blacktip grouper (''Epinephelus fasciatus''), also known as the redbanded grouper, blacktipped cod, black-tipped rockcod, footballer cod, red-barred cod, red-barred rockcod, scarlet rock-cod or weathered rock-cod, is a species of marine
ray-finned fish
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species.
The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
, a
grouper
Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes.
Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...
from the
subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Epinephelinae
Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes.
Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is u ...
which is part of the
family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Serranidae
The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some case ...
, which also includes the
anthias
Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...
and sea basses. It is found in the tropical
Indo-Pacific
The Indo-Pacific is a vast biogeographic region of Earth.
In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region. It is the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Epinephelus
''Epinephelus'' is a genus of marine ray-finned fish, groupers from the subfamily Epinephelinae, part of the family Serranidae, which also includes the anthias and sea basses. They are predatory fish, largely associated with reefs and are found i ...
''.
Description
The blacktip grouper has a body which has a
standard length
Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.
Overall length
* Standard length (SL) is the length of a fish m ...
which is around 2.8 to 3.3 times its depth. The area between the yes is flat but the dorsal profile of the head is convex. The rounded preopercle has a finely serrated rear margin with he lowest serrations slightly enlarged. The upper edge of the gill cover is straight.
The
dorsal fin
A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
contains 11 spines and 15-17 soft rays while the
anal fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has 3 spines and 8 soft rays.
[ The membranes between the dorsal fin spines are deeply indented. The ]caudal fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is moderately rounded. There are 49-75 scales in the lateral line
The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
.[ The colour is variable and ranges from pale greenish grey to pale reddish yellow to scarlet. They frequently have 5 or 6 faint dark bars, the final one being on the caudal peduncle. The scales on the upper body have a pale centre and dark rear margin, which creates am indistinct checked pattern. The outer membrane of the spiny part of the dorsal fin is black, or dark red in specimens from Western Australia and some from deep water. There is a pale yellow or white spot to the rear of the tip of each of the dorsal fin spines. This species attains a maximum ]total length
Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology.
Overall length
* Standard length (SL) is the length of a fish ...
of , although a more common length is around , and a weight of .[
]
Distribution
The blacktip grouper has a wide Indo-Pacific distribution. Its range extends from the Red Sea
The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
to the Eastern Cape
The Eastern Cape is one of the provinces of South Africa. Its capital is Bhisho, but its two largest cities are East London and Gqeberha.
The second largest province in the country (at 168,966 km2) after Northern Cape, it was formed in ...
in South Africa east as far as the Pitcairn Islands
The Pitcairn Islands (; Pitkern: '), officially the Pitcairn, Henderson, Ducie and Oeno Islands, is a group of four volcanic islands in the southern Pacific Ocean that form the sole British Overseas Territory in the Pacific Ocean. The four isl ...
, north to southern Japan and Korea and south to New Caledonia
)
, anthem = ""
, image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg
, map_alt = Location of New Caledonia
, map_caption = Location of New Caledonia
, mapsize = 290px
, subdivision_type = Sovereign st ...
and Australia. It is found in the Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, Mascarenes
The Mascarene Islands (, ) or Mascarenes or Mascarenhas Archipelago is a group of islands in the Indian Ocean east of Madagascar consisting of the islands belonging to the Republic of Mauritius as well as the French department of Réunion. Their ...
, Comoros
The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
and the Seychelles
Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, V ...
in the Indian Ocean as well. In Australia it occurs from the Houtman Abrolhos
The Houtman Abrolhos (often called the Abrolhos Islands) is a chain of 122 islands and associated coral reefs, in the Indian Ocean off the west coast of Australia, about west of Geraldton, Western Australia. It is the southernmost true coral ...
in Western Australia
Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
north around the tropical coastline and then as far south as Port Hacking
Port Hacking Estuary ( Aboriginal Tharawal language: ''Deeban''), an open youthful tide dominated, drowned valley estuary, is located in southern Sydney, New South Wales, Australia approximately south of Sydney central business district. Port ...
in New South Wales
)
, nickname =
, image_map = New South Wales in Australia.svg
, map_caption = Location of New South Wales in AustraliaCoordinates:
, subdivision_type = Country
, subdivision_name = Australia
, established_title = Before federation
, es ...
. It can also be found on reefs in the Coral Sea
The Coral Sea () is a marginal sea of the South Pacific off the northeast coast of Australia, and classified as an interim Australian bioregion. The Coral Sea extends down the Australian northeast coast. Most of it is protected by the Fre ...
, Elizabeth Reef
Elizabeth Reef, located at is a coral reef in the Coral Sea. The reef is separated by a deep oceanic pass, some 47 km wide, from nearby Middleton Reef, both of which are part of the underwater plateau known as the Lord Howe Rise. It i ...
, around Lord Howe Island
Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland P ...
in the Tasman Sea
The Tasman Sea (Māori: ''Te Tai-o-Rēhua'', ) is a marginal sea of the South Pacific Ocean, situated between Australia and New Zealand. It measures about across and about from north to south. The sea was named after the Dutch explorer Abe ...
and Christmas Island
Christmas Island, officially the Territory of Christmas Island, is an Australian external territory comprising the island of the same name. It is located in the Indian Ocean, around south of Java and Sumatra and around north-west of the ...
. A single record was reported in 2012 from the eastern Mediterranean Sea
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the ea ...
, off Lebanon.[Atlas of Exotic Fishes in the Mediterranean Sea (Epinephelus fasciatus). 2nd Edition. 2021. 366p. CIESM Publishers, Paris, Monaco.https://ciesm.org/atlas/fishes_2nd_edition/Epinephelus_fasciatus.pdf]
Biology
This species may present simultaneous hermaphroditism
Simultaneous hermaphroditism is one of the two types of Hermaphrodite, hermaphroditism, the other type being sequential hermaphroditism. In this form of hermaphroditism an individual has Sex organ, sex organs of both Sex, sexes and can produce bo ...
in smaller individuals, while the large individuals usually lose female function.[
The blacktip grouper feeds on ]crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s and smaller fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
es by ambush
An ambush is a long-established military tactics, military tactic in which a combatant uses an advantage of concealment or the element of surprise to attack unsuspecting enemy combatants from concealed positions, such as among dense underbru ...
ing them. It is found associated with coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups.
Co ...
s from 4 m deep (more commonly from 15 m) up to 160 m, in both marine
Marine is an adjective meaning of or pertaining to the sea or ocean.
Marine or marines may refer to:
Ocean
* Maritime (disambiguation)
* Marine art
* Marine biology
* Marine debris
* Marine habitats
* Marine life
* Marine pollution
Military
* ...
and brackish
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water, sometimes in groups of 10-15 individuals. Juveniles may find shelter in mangrove
A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
swamps.
Blacktip groupers of the Red Sea
The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
are fished by the Bedouin
The Bedouin, Beduin, or Bedu (; , singular ) are nomadic Arab tribes who have historically inhabited the desert regions in the Arabian Peninsula, North Africa, the Levant, and Mesopotamia. The Bedouin originated in the Syrian Desert and A ...
. It has also been associated with ciguatera
Ciguatera fish poisoning (CFP), also known simply as ciguatera, is a foodborne illness caused by eating reef fish whose flesh is contaminated with certain toxins. Such individual fish are said to be ciguatoxic. Symptoms may include diarrhea, vomi ...
poisoning.[
]
Parasites
Blacktip groupers are host of several parasite
Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s, including ''Pseudorhabdosynochus
''Pseudorhabdosynochus'' is a genus of monopisthocotylean monogeneans, included in the family Diplectanidae. The type-species of the genus is ''Pseudorhabdosynochus epinepheli'' ( Yamaguti, 1938).
The genus includes more than 80 species, which a ...
'' spp. ( diplectanid Monogenea
Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female reprod ...
ns) on the gills. The philometrid nematode
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhab ...
''Philometra fasciati
''Philometra fasciati'' is a species of parasitic nematode of fishes, first found off New Caledonia in the South Pacific, in the gonads '' Epinephelus fasciatus''. This species is characterized mainly by: length of spicules and length and stru ...
'' is parasitic in the ovary of female fish;[ ] the adult female parasite is a red worm which can reach up to 40 centimetres in length, for a diameter of only 1.6 millimetre; the males are tiny.
''Raphidascaris (Ichthyascaris) fasciati'' is a nematode
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhab ...
parasitic in the intestine, 20 mm in length, described in 2020 and named after the fish.[ ]
Taxonomy
The blacktip grouper was first formally described as ''Perca fasciata'' in 1775 by the Swedish
Swedish or ' may refer to:
Anything from or related to Sweden, a country in Northern Europe. Or, specifically:
* Swedish language, a North Germanic language spoken primarily in Sweden and Finland
** Swedish alphabet, the official alphabet used by ...
speaking Finnish
Finnish may refer to:
* Something or someone from, or related to Finland
* Culture of Finland
* Finnish people or Finns, the primary ethnic group in Finland
* Finnish language, the national language of the Finnish people
* Finnish cuisine
See also ...
born explorer Peter Forsskål
Peter Forsskål, sometimes spelled Pehr Forsskål, Peter Forskaol, Petrus Forskål or Pehr Forsskåhl (11 January 1732 – 11 July 1763) was a Swedish-speaking Finnish explorer, orientalist, naturalist, and an apostle of Carl Linnaeus.
Earl ...
(1732-1763) with the type locality given as Ras Muhammad in the southern Sinai Peninsula
The Sinai Peninsula, or simply Sinai (now usually ) (, , cop, Ⲥⲓⲛⲁ), is a peninsula in Egypt, and the only part of the country located in Asia. It is between the Mediterranean Sea to the north and the Red Sea to the south, and is a l ...
of Egypt. The German
German(s) may refer to:
* Germany (of or related to)
**Germania (historical use)
* Germans, citizens of Germany, people of German ancestry, or native speakers of the German language
** For citizens of Germany, see also German nationality law
**Ger ...
naturalist Marcus Elieser Bloch
Marcus Elieser Bloch (1723–1799) was a German physician and naturalist who is best known for his contribution to ichthyology through his multi-volume catalog of plates illustrating the fishes of the world. Brought up in a Hebrew-speaking Jewish ...
(1723–1799) created the new genus ''Epinephelus'' when he described ''E. marginalis'' in 1793, however ''E. marginalis'' is a synonym
A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of ''Perca fasciata'' and this means that this species is the type species of its genus.
Gallery
File:Epinephelus fasciatus, juvénile.jpg, Juvenile
Epinephelus fasciatus.Групер (Lat. Serranidae). DSCF2701ОВ.jpg, Adult
File: Serranidae - Epinephelus fasciatus.JPG, A variant of an adult with a pale body and a red front part
File:Parasite140007-fig7 Philometra fasciati Moravec & Justine, 2008 (Nematoda, Philometridae).tif, ''Philometra fasciati
''Philometra fasciati'' is a species of parasitic nematode of fishes, first found off New Caledonia in the South Pacific, in the gonads '' Epinephelus fasciatus''. This species is characterized mainly by: length of spicules and length and stru ...
'' (Nematoda
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broa ...
, Philometridae), a parasite of the ovary of the blacktip grouper
References
External links
Wildscreen Archive
*
{{Taxonbar, from=Q2281131
blacktip grouper
Fish of the Red Sea
Fish of the Pacific Ocean
blacktip grouper