Blackburnian Warbler
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The Blackburnian warbler (''Setophaga fusca'') is a small
New World warbler The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. They are not closely related to Old World warblers or Australian warblers. Most ...
. They breed in eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, from southern
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, westwards to the southern
Canadian Prairies The Canadian Prairies (usually referred to as simply the Prairies in Canada) is a region in Western Canada. It includes the Canadian portion of the Great Plains and the Prairie Provinces, namely Alberta, Saskatchewan, and Manitoba. These provin ...
, the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
region and
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
, to
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
. Blackburnian warblers are migratory, wintering in southern
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, and are very rare vagrants to western
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.


Etymology

The genus name ''Setophaga'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''ses'', "moth", and ', "eating", and the specific ''fusca'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for brown. These birds were named after
Anna Blackburne Anna Blackburne (1726 – 30 December 1793) was an English people, English naturalist. Life Anna Blackburne was born at Orford Hall, Orford, Warrington, Lancashire, the daughter of John Blackburne (botanist), John Blackburne and Jane (born Ashto ...
, an
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
.


Description

Blackburnian warblers are small passerines and average-sized wood-warblers. They measure around long, with a wingspan, and weigh . The average mass of an adult bird is , although is slightly higher in fall due to fat reserves, averaging . Among standard measurements, the wing chord is , the
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...
is , the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is and the tarsus is . In summer, male Blackburnian warblers display dark gray backs and double white wing bars, with yellowish rumps and dark brown crowns. The underparts of these birds are white, and are tinged with yellow and streaked black. The head is strongly patterned in yellow and black, with a flaming-orange throat. It is the only North American warbler with this striking plumage. Other plumages, including the fall male and adult female, are washed-out versions of the summer male, and in particular lack the bright colors and strong head pattern. The Blackburnian warbler is practically unmistakable if seen well, even the female due her dull-yellow supercilium, contrasting with greyish cheeks and yellow throat contrasting with the dark streaky sides and back. The only other wood-warbler with an orange throat is the
flame-throated warbler The flame-throated warbler (''Oreothlypis gutturalis'') is a small New World warbler. It is sometimes placed in the genera ''Vermivora'' and ''Parula''. Description The adult flame-throated warbler is long and weighs . It has slate grey upper p ...
of
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and is very distinctive, lacking the contrasting blackish streaking about the head and whitish underside of a male Blackburnian. Basic plumages show weaker yellows and gray in place of black in the breeding male. Blackburnian warblers' songs are a simple series of high ''swi'' notes, which often ascend in pitch. Transliterations have included ''zip zip zip zip zip zip zip zip'', ''titititi tseeeeee'' or ''teetsa teetsa teetsa teetsa''. Their call is a high ''sip''. Genetic research has shown that their closed living relative is the
bay-breasted warbler The bay-breasted warbler (''Setophaga castanea'') is a small species of songbird in the New World warbler family, Parulidae. It is one of thirty-four species in the diverse genus ''Setophaga.'' Like all songbirds, or passerines, the species is cl ...
, the latter species perhaps specialized to forage in the same coniferous trees at lower levels. Hybridization in the wild has been recorded once each with a bay-breasted warbler (in
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...
), with a
black-and-white warbler The black-and-white warbler (''Mniotilta varia'') is a species of New World warbler, and the only member of its genus, ''Mniotilta''. It breeds in northern and eastern North America and winters in Florida, Central America, and the West Indies ...
(in
Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
) and possibly a wintering hybrid with a
Kirtland's warbler Kirtland's warbler (''Setophaga kirtlandii''), also known in Michigan by the common name jack pine bird, or the jack pine warbler, is a small songbird of the New World warbler family (Parulidae), named after Jared Potter Kirtland, an Ohio doctor ...
(in
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
).


Ecology

Blackburnian warblers are solitary during winter and highly territorial on their breeding grounds and do not mix with other passerine species outside of the migratory period. However, during migration, they often join local mixed foraging flocks of species such as
chickadee The chickadees are a group of North American birds in the tit family included in the genus ''Poecile''. Species found in North America are referred to as chickadees, while other species in the genus are called tits. They are small-sized birds ...
s,
kinglet A kinglet is a small bird in the family Regulidae. Species in this family were formerly classified with the Old World warblers. "Regulidae" is derived from the Latin word ''regulus'' for "petty king" or prince, and refers to the coloured crowns ...
s and
nuthatch The nuthatches () constitute a genus, ''Sitta'', of small passerine birds belonging to the family Sittidae. Characterised by large heads, short tails, and powerful bills and feet, nuthatches advertise their territory using loud, simple songs. Mo ...
es. Similarly, in the tropics they were found to be fairly social while engaging in migration but solitary from other passerines while wintering.Bent, A. C. 1953. ''Life histories of North American wood warblers''. U.S. Natl. Mus. Bull. 203. These birds are basically insectivorous, but will include berries in their diets in wintertime. They usually forage by searching for insects or
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s in treetops. Their breeding season diet is dominated by the larvae of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
, i.e. moths and butterflies. They may help control the
spruce budworm ''Choristoneura'' is a genus of moths in the family Tortricidae. Several species are serious pests of conifers, such as spruce and are known as spruce budworms. Species *''Choristoneura adumbratanus'' (Walsingham, 1900) *'' Choristoneura afric ...
(often considered a harmful pest) when breakouts occur, at the local if not at epidemic level. In one study from
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
, 98% of the diet was made of insects, the remaining 2% being spiders. Among the migratory ''Setophaga'' warblers, it is considered one of the specialist at foraging in the micro-habitat of the tree's top canopy. The breeding habitats of these birds are mature
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous woodlands, the central part of their breeding range being in the southeastern portion of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
's
boreal forest Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, ...
. However, their distribution as a breeding species continues broadly down much of
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
and the
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
, from
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...
to northernmost
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, in elevated mixed woodlands, especially ones containing
spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
and hemlocks.Morse, Douglass H. 2004. ''Blackburnian Warbler (Setophaga fusca)'', The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. Hemlocks in particular are most likely to host Blackburnian warblers in mixed forests. It typically winters in tropical montane forests, from roughly , mainly from
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
to
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
, more sporadically in
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
and the
Amazon region The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
. Blackburnian warblers begin their first clutches in mid-May to early June in the contiguous United States and about 1 to 2 weeks later in
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
. This species build a nest consisting of an open cup of twigs, bark, plant fibers, and rootlets held to branch with spider web and lined with lichens, moss, hair, and dead pine needles that's placed near the end of a branch. Although typically only laying one brood per year, if a nest is destroyed they are capable of producing a second or even third brood. Three to five whitish eggs are laid in its nest which is usually placed above the ground, on a horizontal branch. Nests usually constructed outwardly with twigs, bark, plant fibers, and rootlets; lined with lichens, mosses, fine grasses, hair, dead pine needles, and even occasionally such exotic substances as string, willow cotton, horsehair, and cattail down. Only the female broods and spends about 80% day actively brooding, with the male usually helping bring food to the nest. Among warblers, they are relatively rarely parasitized at the nest by
brown-headed cowbird The brown-headed cowbird (''Molothrus ater'') is a small, obligate brood parasitic icterid native to temperate and subtropical North America. It is a permanent resident in the southern parts of its range; northern birds migrate to the southern ...
s, most likely due to the cowbirds lack of success in dense pine-dominated forests.
Blue jay The blue jay (''Cyanocitta cristata'') is a passerine bird in the family Corvidae, native to eastern North America. It lives in most of the eastern and central United States; some eastern populations may be migratory. Resident populations are a ...
s and
American red squirrel The American red squirrel (''Tamiasciurus hudsonicus'') is one of three species of tree squirrels currently classified in the genus ''Tamiasciurus'', known as the pine squirrels (the others are the Douglas squirrel, ''T. douglasii'', and the sou ...
s have been verified to prey on nestlings and new fledglings, while a
merlin Merlin ( cy, Myrddin, kw, Marzhin, br, Merzhin) is a mythical figure prominently featured in the legend of King Arthur and best known as a mage, with several other main roles. His usual depiction, based on an amalgamation of historic and le ...
was recorded killing a brooding adult female.
Sharp-shinned hawk The sharp-shinned hawk (''Accipiter striatus'') is a small hawk, with males being the smallest hawks in the United States and Canada, but with the species averaging larger than some Neotropical realm, Neotropical species, such as the tiny hawk. ...
s and
Cooper's hawk Cooper's hawk (''Accipiter cooperii'') is a medium-sized hawk native to the North American continent and found from southern Canada to Mexico. This species is a member of the genus ''Accipiter'', sometimes referred to as true hawks, which are f ...
s are likely, but not confirmed, predators of adult Blackburnian warblers. By far the greatest threat faced by this species is destruction of forest habitat, which some predict could cause the Blackburnian warbler to lose up more than 30% of its wintering or breeding habitat.Diamond, A. W. 1991. ''Assessment of the risks from tropical deforestation to Canadian songbirds''. Trans. NA Wildl. Nat. Res. Conf. 56:177-194. However, currently this species continues to occur over a large range and can appear in stable numbers where habitat is appropriate.


References


External links

*
Blackburnian warbler species profile
- The Nature Conservancy

- Cornell Lab of Ornithology

- USGS Patuxent Bird Identification InfoCenter

(for
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
,
Grenada Grenada ( ; Grenadian Creole French: ) is an island country in the West Indies in the Caribbean Sea at the southern end of the Grenadines island chain. Grenada consists of the island of Grenada itself, two smaller islands, Carriacou and Pe ...
) with Range Map at bird-stamps.org * {{Taxonbar, from=Q27075919
Blackburnian warbler The Blackburnian warbler (''Setophaga fusca'') is a small New World warbler. They breed in eastern North America, from southern Canada, westwards to the southern Canadian Prairies, the Great Lakes region and New England, to North Carolina. ...
Native birds of Eastern Canada Native birds of the Northeastern United States Birds of Appalachia (United States) Birds of Cuba Birds of Grenada Birds of South America
Blackburnian warbler The Blackburnian warbler (''Setophaga fusca'') is a small New World warbler. They breed in eastern North America, from southern Canada, westwards to the southern Canadian Prairies, the Great Lakes region and New England, to North Carolina. ...
Blackburnian warbler The Blackburnian warbler (''Setophaga fusca'') is a small New World warbler. They breed in eastern North America, from southern Canada, westwards to the southern Canadian Prairies, the Great Lakes region and New England, to North Carolina. ...