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The blackberry is an edible
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
produced by many
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
in the genus ''
Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species. Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of these ...
'' in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...
, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of blackberries has historically been confused because of
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
and
apomixis In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cuttin ...
, so that species have often been grouped together and called
species aggregate In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
s. For example, the entire subgenus ''Rubus'' has been called the ''
Rubus fruticosus ''Rubus fruticosus'' L. is the ambiguous name of a European blackberry species in the genus ''Rubus'' in the rose family. The name has been interpreted in several ways: *The species represented by the type specimen of ''Rubus fruticosus'' L., ...
'' aggregate, although the species ''R. fruticosus'' is considered a synonym of '' R. plicatus''. ''
Rubus armeniacus ''Rubus armeniacus'', the Himalayan blackberry or Armenian blackberry, is a species of ''Rubus'' in the blackberry group ''Rubus'' subgenus ''Rubus'' series ''Discolores'' (P.J. Müll.) Focke. It is native to Armenia and Northern Iran, and wide ...
'' ("Himalayan" blackberry) is considered a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is injurious to agricultural or horticultural crops, natural habitats or ecosystems, or humans or liv ...
and
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
in many regions of the
Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though ...
of Canada and the United States, where it grows out of control in urban and suburban parks and woodlands.


Description

What distinguishes the blackberry from its
raspberry The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...
relatives is whether or not the torus ( receptacle or stem) "picks with" (i.e., stays with) the
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
. When picking a blackberry fruit, the torus stays with the fruit. With a raspberry, the torus remains on the plant, leaving a hollow core in the raspberry fruit. The term ''
bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...
'', a word referring to any impenetrable
thicket A thicket is a very dense stand of trees or tall shrubs, often dominated by only one or a few species, to the exclusion of all others. They may be formed by species that shed large numbers of highly viable seeds that are able to germinate in the ...
, has in some circles traditionally been applied specifically to the blackberry or its products, though in the United States it applies to all members of the genus ''Rubus''. In small parts of the western US, the term ''caneberry'' is used to refer to blackberries and raspberries as a group rather than the term bramble. ''Briar'' or ''brier'' is also sometimes used to refer to the plant, though this name is used for other thorny thickets (such as ''
Smilax ''Smilax'' is a genus of about 300–350 species, found in the tropics and subtropics worldwide. In China for example about 80 are found (39 of which are endemic), while there are 20 in North America north of Mexico. They are climbing flowering ...
'') as well. The usually black fruit is not a
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
in the
botanical Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
sense of the word. Botanically it is termed an
aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...
, composed of small
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
lets. It is a widespread and well-known group of over 375 species, many of which are closely related apomictic microspecies
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
throughout Europe, northwestern Africa,
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
western and central Asia and North and South America.


Botanical characteristics

Blackberries are
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s which typically bear
biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...
stems ("canes") from the perennial root system. In its first year, a new stem, the primocane, grows vigorously to its full length of (in some cases, up to ), arching or trailing along the ground and bearing large palmately compound
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
with five or seven leaflets; it does not produce any flowers. In its second year, the cane becomes a floricane and the stem does not grow longer, but the lateral buds break to produce flowering laterals (which have smaller leaves with three or five leaflets). First- and second-year shoots usually have numerous short-curved, very sharp prickles that are often erroneously called thorns. These prickles can tear through denim with ease and make the plant very difficult to navigate around. Prickle-free cultivars have been developed. The
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a public land-grant research university in Fayetteville, Arkansas. It is the flagship campus of the University of Arkansas System and the largest university in the state. Founded as Arkansas ...
has developed primocane-fruiting blackberries that grow and flower on first-year growth as much as the primocane-fruiting (also called fall bearing or everbearing) red
raspberries The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...
do. Unmanaged mature plants form a tangle of dense arching stems, the branches rooting from the node tip on many species when they reach the ground. Vigorous and growing rapidly in woods, scrub, hillsides, and hedgerows, blackberry shrubs tolerate poor
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former te ...
s, readily colonizing wasteland, ditches, and vacant lots. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are produced in late spring and early summer on short
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s on the tips of the flowering laterals. Each flower is about in diameter, with five white or pale pink
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s. The drupelets only develop around ovules that are fertilized by the male gamete from a pollen grain. The most likely cause of undeveloped ovules is inadequate
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
visits. Even a small change in conditions, such as a rainy day or a day too hot for bees to work after early morning, can reduce the number of bee visits to the flower, thus reducing the quality of the fruit. Incomplete drupelet development can also be a symptom of exhausted reserves in the plant's roots or infection with a
virus A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsky's 1 ...
such as
raspberry bushy dwarf virus ''Idaeovirus'' is a genus of positive-sense ssRNA viruses that contains two species: ''Raspberry bushy dwarf virus'' (RBDV) and ''Privet idaeovirus''. RBDV has two host-dependent clades: one for raspberries; the other for grapevines. Infection ...
.


History

One of the earliest known instances of blackberry consumption comes from the remains of the
Haraldskær Woman The Haraldskær Woman (or Haraldskjaer Woman) is the name given to a bog body of a woman preserved in a bog in Jutland, Denmark, and dating from about 490 BC (pre-Roman Iron Age). Workers found the body in 1835 while excavating peat on the Harald ...
, the naturally preserved
bog body A bog body is a human cadaver that has been naturally mummified in a peat bog. Such bodies, sometimes known as bog people, are both geographically and chronologically widespread, having been dated to between and the Second World War. Fischer 199 ...
of a
Danish Danish may refer to: * Something of, from, or related to the country of Denmark People * A national or citizen of Denmark, also called a "Dane," see Demographics of Denmark * Culture of Denmark * Danish people or Danes, people with a Danish ance ...
woman dating from approximately 2,500 years ago. Forensic evidence found blackberries in her stomach contents, among other foods. The use of blackberries to make wines and cordials was documented in the
London Pharmacopoeia A pharmacopoeia, pharmacopeia, or pharmacopoea (from the obsolete typography ''pharmacopœia'', meaning "drug-making"), in its modern technical sense, is a book containing directions for the identification of compound medicines, and published by ...
in 1696. In the culinary world, blackberries have a long history of use alongside other fruits to make pies, jellies and jams. Blackberry plants were used for
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
by Greeks, other European peoples, and aboriginal Americans. A 1771 document described brewing blackberry leaves, stem, and bark for
stomach ulcers Peptic ulcer disease (PUD) is a break in the inner lining of the stomach, the first part of the small intestine, or sometimes the lower esophagus. An ulcer in the stomach is called a gastric ulcer, while one in the first part of the intestines ...
. Blackberry fruit, leaves, and stems have been used to dye fabrics and hair. Native Americans have even been known to use the stems to make rope. The shrubs have also been used for barriers around buildings, crops and livestock. The wild plants have sharp, thick prickles, which offered some protection against enemies and large animals.


Cultivar development

Modern
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
and
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
development took place mostly in the United States. In 1880, a hybrid blackberry-raspberry named the
loganberry The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry (''Rubus ursinus'') and the European raspberry (''Rubus idaeus''). The plant and the fruit resemble the blackberry more than the raspberry, but the frui ...
was developed in
Santa Cruz, California Santa Cruz (Spanish for "Holy Cross") is the county seat and largest city of Santa Cruz County, in Northern California. As of the 2020 census, the city population was 62,956. Situated on the northern edge of Monterey Bay, Santa Cruz is a pop ...
, by an American judge and
horticulturalist Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
,
James Harvey Logan James Harvey Logan (December 8, 1841 - July 16, 1928) was a judge in Santa Cruz, California, and an amateur botanist credited with the 1881 creation of the loganberry, a cross between the raspberry and the blackberry. He was District Attorney i ...
. One of the first thornless varieties was developed in 1921, but the berries lost much of their flavor. Common thornless cultivars developed from the 1990s to the early 21st century by the
US Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
enabled efficient machine-harvesting, higher yields, larger and firmer fruit, and improved flavor, including the ''Triple Crown'', ''Black Diamond'', ''Black Pearl'', and ''Nightfall'', a
marionberry The marionberry (''Rubus L.'' subgenus ''Rubus'') is a cultivar of blackberry developed by the USDA ARS breeding program in cooperation with Oregon State University. A cross between the ' Chehalem' and ' Olallie' varieties,
.


Ecology

Blackberry leaves are food for certain
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s; some grazing mammals, especially deer, are also very fond of the leaves. Caterpillars of the
concealer moth Oecophoridae (concealer moths) is a family of small moths in the superfamily Gelechioidea. The phylogeny and systematics of gelechoid moths are still not fully resolved, and the circumscription of the Oecophoridae is strongly affected by this. ...
''
Alabonia geoffrella ''Alabonia geoffrella'' is a species of gelechioid moth. Here, it is placed within the subfamily Oecophorinae of the concealer moth family (Oecophoridae). Alternatively it has been placed in the Elachistidae or Depressariinae together with its p ...
'' have been found feeding inside dead blackberry shoots. When mature, the berries are eaten and their seeds dispersed by mammals, such as the red fox, American black bear and the Eurasian badger, as well as by small birds. Blackberries grow wild throughout most of Europe. They are an important element in the ecology of many countries, and harvesting the berries is a popular pastime. However, their rigorous growth and tendency to grow unchecked if not managed correctly means that the plants are also considered a
weed A weed is a plant considered undesirable in a particular situation, "a plant in the wrong place", or a plant growing where it is not wanted.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. ...
, sending down roots from branches that touch the ground, and sending up suckers from the roots. In some parts of the world, such as in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, and the Pacific Northwest of North America, some blackberry species, particularly ''
Rubus armeniacus ''Rubus armeniacus'', the Himalayan blackberry or Armenian blackberry, is a species of ''Rubus'' in the blackberry group ''Rubus'' subgenus ''Rubus'' series ''Discolores'' (P.J. Müll.) Focke. It is native to Armenia and Northern Iran, and wide ...
'' (Himalayan blackberry) and ''
Rubus laciniatus ''Rubus laciniatus'', the cutleaf evergreen blackberry or evergreen blackberry, is a species of ''Rubus'', native to Eurasia. It is an introduced species in Australia and North America. It has become a weed and invasive species in forested habita ...
'' (evergreen blackberry), are
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...
and considered an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
and a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is injurious to agricultural or horticultural crops, natural habitats or ecosystems, or humans or liv ...
. Blackberry fruits are red when unripe, leading to an old expression that "blackberries are red when they're green".


Uses


Nutrients

Raw blackberries are 88% water, 10%
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s, 1%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
, and 0.5%
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers spec ...
(table). In a reference amount, raw cultivated blackberries supply 43
calorie The calorie is a unit of energy. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of on ...
s and rich contents (20% or more of the
Daily Value The Reference Daily Intake (RDI) used in nutrition labeling on food and dietary supplement products in the U.S. and Canada is the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97–98% of health ...
(DV) of
dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...
,
manganese Manganese is a chemical element with the symbol Mn and atomic number 25. It is a hard, brittle, silvery metal, often found in minerals in combination with iron. Manganese is a transition metal with a multifaceted array of industrial alloy use ...
(31% DV),
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) an ...
(25% DV), and
vitamin K Vitamin K refers to structurally similar, fat-soluble vitamers found in foods and marketed as dietary supplements. The human body requires vitamin K for post-synthesis modification of certain proteins that are required for blood coagulation ...
(19% DV) (table).


Seed composition

Blackberries contain numerous large seeds that are not always preferred by consumers. The seeds contain oil rich in
omega-3 Omega−3 fatty acids, also called Omega-3 oils, ω−3 fatty acids or ''n''−3 fatty acids, are polyunsaturated fatty acids (PUFAs) characterized by the presence of a double bond, three atoms away from the terminal methyl group in their chem ...
(
alpha-linolenic acid ''alpha''-Linolenic acid (ALA), also known as α-Linolenic acid (from Greek ''alpha'' meaning "first" and ''linon'' meaning flax), is an ''n''−3, or omega-3, essential fatty acid. ALA is found in many seeds and oils, including flaxseed, waln ...
) and omega-6 (
linoleic acid Linoleic acid (LA) is an organic compound with the formula COOH(CH2)7CH=CHCH2CH=CH(CH2)4CH3. Both alkene groups are cis-trans isomerism, ''cis''. It is a fatty acid sometimes denoted 18:2 (n-6) or 18:2 ''cis''-9,12. A linoleate is a salt (chem ...
) fats as well as
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
, dietary fiber,
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic compound, organic pigments that are produced by plants and algae, as well as several bacteria, and Fungus, fungi. Carotenoids give the characteristic color to pumpki ...
s,
ellagitannins The ellagitannins are a diverse class of hydrolyzable tannins, a type of polyphenol formed primarily from the oxidative linkage of galloyl groups in 1,2,3,4,6-pentagalloyl glucose. Ellagitannins differ from gallotannins, in that their galloyl group ...
, and
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backwards because it can be ob ...
.


Culinary use

The ripe fruit is commonly used in
dessert Dessert is a course (food), course that concludes a meal. The course consists of sweet foods, such as confections, and possibly a beverage such as dessert wine and liqueur. In some parts of the world, such as much of Greece and West Africa, and ...
s, jams, jelly, wine and
liqueur A liqueur (; ; ) is an alcoholic drink composed of spirits (often rectified spirit) and additional flavorings such as sugar, fruits, herbs, and spices. Often served with or after dessert, they are typically heavily sweetened and un-aged beyond ...
s. It may be mixed with other berries and fruits for pies and
crumble A crumble is a dish that can be made in a sweet or savoury version. Crumbles became popular in Britain during World War II, when the topping was an economical alternative to pies due to shortages of pastry ingredients as the result of rationi ...
s. Blackberries are also used to produce candy.


Phytochemical research

Blackberries contain numerous
phytochemicals Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poison ...
including
polyphenols Polyphenols () are a large family of naturally occurring organic compounds characterized by multiples of phenol units. They are abundant in plants and structurally diverse. Polyphenols include flavonoids, tannic acid, and ellagitannin, some of ...
,
flavonoids Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
,
anthocyanin Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...
s,
salicylic acid Salicylic acid is an organic compound with the formula HOC6H4CO2H. A colorless, bitter-tasting solid, it is a precursor to and a metabolite of aspirin (acetylsalicylic acid). It is a plant hormone, and has been listed by the EPA Toxic Substance ...
,
ellagic acid Ellagic acid is a polyphenol found in numerous fruits and vegetables. It is the dilactone of hexahydroxydiphenic acid. Name The name comes from the French term ''acide ellagique'', from the word ''galle'' spelled backwards because it can be ob ...
, and
fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
.
Anthocyanins Anthocyanins (), also called anthocyans, are water-soluble vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart gave the name Anthokyan to a chemical compo ...
in blackberries are responsible for their rich dark color. One report placed blackberries at the top of more than 1,000 polyphenol-rich foods consumed in the United States, but this concept of a health benefit from consuming darkly colored foods like blackberries remains scientifically unverified and not accepted for
health claim A health claim on a food label and in food marketing is a claim by a manufacturer of food products that their food will reduce the risk of developing a disease or condition. For example, it is claimed by the manufacturers of oat cereals that oat ...
s on food labels.


Cultivation

Worldwide, Mexico is the leading producer of blackberries, with nearly the entire crop being produced for export into the off-season fresh markets in North America and Europe. Until 2018, the Mexican market was almost entirely based on the cultivar 'Tupy' (often spelled 'Tupi', but the EMBRAPA program in Brazil from which it was released prefers the 'Tupy' spelling), but Tupy fell out of favor in some Mexican growing regions. In the US,
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
is the leading commercial blackberry producer, producing on in 2017. Numerous
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...
s have been selected for commercial and amateur cultivation in Europe and the United States. Since the many species form
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
s easily, there are numerous cultivars with more than one species in their ancestry.


Hybrids

'Marion' (marketed as "
marionberry The marionberry (''Rubus L.'' subgenus ''Rubus'') is a cultivar of blackberry developed by the USDA ARS breeding program in cooperation with Oregon State University. A cross between the ' Chehalem' and ' Olallie' varieties,
") is an important cultivar that was selected from seedlings from a cross between 'Chehalem' and 'Olallie' (commonly called "
Olallieberry The olallieberry ( ), sometimes spelled ollalieberry, olallaberry, olalliberry, ollalaberry or ollaliberry, is the marketing name for the 'Olallie' blackberry released by the USDA-ARS (in collaboration with Oregon State University). The berry was ...
") berries. 'Olallie' in turn is a cross between
loganberry The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry (''Rubus ursinus'') and the European raspberry (''Rubus idaeus''). The plant and the fruit resemble the blackberry more than the raspberry, but the frui ...
and
youngberry The youngberry is a complex hybrid between three different berry species from the genus ''Rubus'' of the rose family: raspberry, blackberry, and dewberry. The berries of the plant are eaten fresh or used to make juice, jam, and in recipes. The y ...
. 'Marion', 'Chehalem' and 'Olallie' are just three of many trailing blackberry cultivars developed by the
United States Department of Agriculture The United States Department of Agriculture (USDA) is the United States federal executive departments, federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, ...
Agricultural Research Service The Agricultural Research Service (ARS) is the principal in-house research agency of the United States Department of Agriculture (USDA). ARS is one of four agencies in USDA's Research, Education and Economics mission area. ARS is charged with ext ...
(USDA-ARS) blackberry breeding program at
Oregon State University Oregon State University (OSU) is a public land-grant, research university in Corvallis, Oregon. OSU offers more than 200 undergraduate-degree programs along with a variety of graduate and doctoral degrees. It has the 10th largest engineering co ...
in
Corvallis, Oregon Corvallis ( ) is a city and the county seat of Benton County in central western Oregon, United States. It is the principal city of the Corvallis, Oregon Metropolitan Statistical Area, which encompasses all of Benton County. As of the 2020 United ...
. The most recent cultivars released from this program are the prickle-free cultivars 'Black Diamond', 'Black Pearl', and 'Nightfall' as well as the very early-ripening 'Obsidian' and 'Metolius'. 'Black Diamond' is now the leading cultivar being planted in the Pacific Northwest. Some of the other cultivars from this program are 'Newberry', 'Waldo', 'Siskiyou', 'Black Butte', ' Kotata', 'Pacific', and 'Cascade'.


Trailing

Trailing blackberries are vigorous and crown forming, require a trellis for support, and are less cold hardy than the erect or semi-erect blackberries. In addition to the
Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though ...
, these types do well in similar climates, such as the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, and the
Mediterranean countries The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the eas ...
.


Thornless

Semi-erect, prickle-free blackberries were first developed at the
John Innes Centre The John Innes Centre (JIC), located in Norwich, Norfolk, England, is an independent centre for research and training in plant and microbial science founded in 1910. It is a registered charity (No 223852) grant-aided by the Biotechnology and B ...
in
Norwich Norwich () is a cathedral city and district of Norfolk, England, of which it is the county town. Norwich is by the River Wensum, about north-east of London, north of Ipswich and east of Peterborough. As the seat of the See of Norwich, with ...
, UK, and subsequently by the USDA-ARS in
Beltsville, Maryland Beltsville is a census-designated place (CDP) in northern Prince George's County, Maryland, United States. The community was named for Truman Belt, a local landowner. The 2020 census counted 20,133 residents. Beltsville includes the unincorporated ...
. These are crown forming and very vigorous and need a trellis for support. Cultivars include 'Black Satin', 'Chester Thornless', 'Dirksen Thornless', 'Hull Thornless', 'Loch Maree', 'Loch Ness', 'Loch Tay', 'Merton Thornless', 'Smoothstem', and 'Triple Crown'. 'Loch Ness' and 'Loch Tay' have gained the RHS's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
. The cultivar 'Cacanska Bestrna' (also called 'Cacak Thornless') has been developed in Serbia and has been planted on many thousands of hectares there.


Erect

The
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a public land-grant research university in Fayetteville, Arkansas. It is the flagship campus of the University of Arkansas System and the largest university in the state. Founded as Arkansas ...
has developed cultivars of erect blackberries. These types are less vigorous than the semi-erect types and produce new canes from root initials (therefore they spread underground like raspberries). There are prickly and prickle-free cultivars from this program, including 'Navaho', 'Ouachita', 'Cherokee', 'Apache', 'Arapaho', and 'Kiowa'. They are also responsible for developing the primocane fruiting blackberries such as 'Prime-Jan' and 'Prime-Jim'.


Primocane

In raspberries, these types are called primocane fruiting, fall fruiting, or everbearing. 'Prime-Jim' and 'Prime-Jan' were released in 2004 by the
University of Arkansas The University of Arkansas (U of A, UArk, or UA) is a public land-grant research university in Fayetteville, Arkansas. It is the flagship campus of the University of Arkansas System and the largest university in the state. Founded as Arkansas ...
and are the first cultivars of primocane fruiting blackberry. They grow much like the other erect cultivars described above; however, the canes that emerge in the spring will flower in midsummer and fruit in late summer or fall. The fall crop has its highest quality when it ripens in cool mild climate such as in California or the Pacific Northwest. 'Illini Hardy', a semi-erect prickly cultivar introduced by the
University of Illinois The University of Illinois Urbana-Champaign (U of I, Illinois, University of Illinois, or UIUC) is a public land-grant research university in Illinois in the twin cities of Champaign and Urbana. It is the flagship institution of the University ...
, is cane hardy in zone 5, where blackberry production has traditionally been problematic, since canes often failed to survive the winter.


Mexico and Chile

Blackberry production in Mexico expanded considerably in the early 21st century. In 2017, Mexico had 97% of the
market share Market share is the percentage of the total revenue or sales in a market that a company's business makes up. For example, if there are 50,000 units sold per year in a given industry, a company whose sales were 5,000 of those units would have a ...
for fresh blackberries imported into the United States, while
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
had 61% of the market share for American imports of frozen blackberries. While once based on the cultivar 'Brazos', an old erect blackberry cultivar developed in Texas in 1959, the Mexican industry is now dominated by the Brazilian 'Tupy' released in the 1990s. The 'Tupy' has the erect blackberry 'Comanche', and a "wild Uruguayan blackberry" as parents. Since there are no native blackberries in Uruguay, the suspicion is that the widely grown '
Boysenberry The boysenberry is a cross between the European raspberry (''Rubus idaeus''), European blackberry (''Rubus fruticosus''), American dewberry (''Rubus aboriginum''), and loganberry (''Rubus'' × ''loganobaccus''). It is a large aggregate fruit, ...
' is the male parent. In order to produce these blackberries in regions of Mexico where there is no winter chilling to stimulate flower bud development, chemical defoliation and application of growth regulators are used to bring the plants into bloom.


Diseases and pests

Because blackberries belong to the same genus as raspberries, they share the same diseases, including
anthracnose A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
, which can cause the berry to have uneven ripening. Sap flow may also be slowed. They also share the same remedies, including the Bordeaux mixture, a combination of lime, water and
copper(II) sulfate Copper(II) sulfate, also known as copper sulphate, is an inorganic compound with the chemical formula . It forms hydrates , where ''n'' can range from 1 to 7. The pentahydrate (''n'' = 5), a bright blue crystal, is the most commonly encountered hy ...
. The rows between blackberry plants must be free of weeds, blackberry suckers and grasses, which may lead to pests or diseases. Fruit growers are selective when planting blackberry bushes because wild blackberries may be infected, and gardeners are recommended to purchase only certified disease-free plants. The spotted-wing drosophila, ''
Drosophila suzukii ''Drosophila suzukii'', commonly called the spotted wing drosophila or SWD, is a fruit fly. ''D. suzukii'', originally from southeast Asia, is becoming a major pest species in America and Europe, because it infests fruit early during the ripening ...
'', is a serious pest of blackberries. Unlike its vinegar fly relatives, which are primarily attracted to rotting or fermented fruit, ''D. suzukii'' attacks fresh, ripe fruit by laying eggs under the soft skin. The larvae hatch and grow in the fruit, destroying the fruit's commercial value. Another pest is ''Amphorophora rubi'', known as the blackberry aphid, which eats not just blackberries but raspberries as well. ''Byturus tomentosus'' (
raspberry beetle The raspberry beetle (''Byturus tomentosus'') is a species of beetle in the fruitworm family Byturidae. It is a major pest that is widespread in north-central Europe, affecting raspberry, blackberry and loganberry plants. This species is rela ...
), ''
Lampronia corticella ''Lampronia corticella'', the raspberry moth, is a moth of the family Prodoxidae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It is found in most of Europe, except Iceland, the Iberian Penin ...
'' (raspberry moth) and ''
Anthonomus rubi ''Anthonomus rubi'', the strawberry-blossom weevil or strawberry blossom weevil, is a species of weevil found in Europe and western Asia. It feeds on plants of the family Rosaceae and is an important pest of strawberry (''Fragaria'' × ''ananassa ...
'' (strawberry blossom weevil) are also known to infest blackberries.


Genetics

The loci controlling the primocane fruiting was mapped in the F Locus, on LG7, whereas thorns/thornlessness was mapped on LG4. Better understanding of the genetics is useful for genetic screening of cross-breds, and for genetic engineering purposes.


Folklore

Folklore in the United Kingdom and Ireland tells that blackberries should not be picked after
Old Michaelmas Day Michaelmas ( ; also known as the Feast of Saints Michael, Gabriel, and Raphael, the Feast of the Archangels, or the Feast of Saint Michael and All Angels) is a Christian festival observed in some Western liturgical calendars on 29 September, ...
(11 October) as the
devil A devil is the personification of evil as it is conceived in various cultures and religious traditions. It is seen as the objectification of a hostile and destructive force. Jeffrey Burton Russell states that the different conceptions of t ...
(or a
Púca The púca (Irish for ''spirit/ghost''; plural púcaí), puca (Old English for ''goblin''; plural pucel) pwca, pooka, phouka, puck is a creature of Celtic, English, and Channel Islands folklore. Considered to be bringers both of good and bad f ...
) has made them unfit to eat by stepping, spitting or fouling on them. There is some value in this legend as autumn's wetter and cooler weather often allows the fruit to become infected by various
mold A mold () or mould () is one of the structures certain fungus, fungi can form. The dust-like, colored appearance of molds is due to the formation of Spore#Fungi, spores containing Secondary metabolite#Fungal secondary metabolites, fungal seco ...
s such as ''
Botryotinia ''Botryotinia'' is a genus of ascomycete fungi causing several plant diseases. The anamorphs of ''Botryotinia'' are mostly included in the " imperfect fungi" genus ''Botrytis''. The genus contains 22 species and one hybrid. Plant diseases caus ...
'' which give the fruit an unpleasant look and may be toxic. According to some traditions, a blackberry's deep purple color represents Christ's blood and the
crown of thorns According to the New Testament, a woven crown of thorns ( or grc, ἀκάνθινος στέφανος, akanthinos stephanos, label=none) was placed on the head of Jesus during the events leading up to his crucifixion. It was one of the instru ...
was made of brambles, although other thorny plants, such as ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' (hawthorn) and ''
Euphorbia milii ''Euphorbia milii'', the crown of thorns, Christ plant, or Christ thorn, is a species of flowering plant in the spurge family Euphorbiaceae, native to Madagascar. The species name commemorates Baron Milius, once Governor of Réunion, who introdu ...
'' (crown of thorns plant), have been proposed as the material for the crown.


See also

*
Mulberry ''Morus'', a genus of flowering plants in the family Moraceae, consists of diverse species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 identif ...
, similar fruit appearance, but a tree rather than a
bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...
* List of Lepidoptera that feed on ''Rubus'' * Redberry mite, a common pest of North American blackberry crops


References


Further reading

*


External links

* {{Commons-inline Berries Medicinal plants Plant common names Rubus