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''Diplocarpon rosae'' is a
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
that creates the
rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...
black spot disease. Because it was observed by people of various countries around the same time (around 1830), the nomenclature for the fungus varied with about 25 different names. The asexual stage is now known to be ''
Marssonina ''Marssonina'' is a genus of fungi in the family Dermateaceae. The genus name of ''Marssonina'' is in honour of Theodor Friedrich Marsson (1816–1892), who was a German pharmacist and botanist Botany, also called , plant biology or ph ...
rosae'', while the sexual and most common stage is known as ''Diplocarpon rosae''. ''Diplocarpon rosae'' grows over seasons as
mycelia Mycelium (plural mycelia) is a root-like structure of a fungus consisting of a mass of branching, thread-like hyphae. Fungal colonies composed of mycelium are found in and on soil and many other substrates. A typical single spore germinates ...
,
ascospore An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or ...
s, and conidia in infected leaves and canes. In the spring during moist, humid conditions, ascospores and conidia are wind-borne and rain-splashed to newly emerging leaf tissue.


Diagnosis

The black spots are circular with a perforated edge, and reach a diameter of . Badly affected plants, however, will not show the circular patterning, as they combine to cause a large, black mass. The common treatment of the disease is to remove the affected leaves and spray with antifungal solutions. Some stems of the roses may become affected if untreated, and will cause progressive weakening of the rose.


Disease cycle

''Diplocarpon rosae'' tends to overwinter in both lesions of infected canes and fallen foliage. Conidia are produced in the diseased stem tissues and dispersed via water—most commonly by rain or wind—into the openings of leaves in the spring season. The conidia then produce germ tubes (and sometimes appressoria) to penetrate the tissues of the leaves. Mycelia develop on the underside of the leaf cuticle and lesions appear. As these lesions appear, acervuli continuously produce conidia asexually as long as the climate remains optimally wet and warm. These conidia can then be dispersed to new uninfected leaves as a source of secondary inoculum, adding more cycles of infection. Once defoliation occurs in the fall season, the hyphae of the ''Diplocarpon'' ''rosae'' invade the dead leaf tissue and form pycnidia lined with conidiophores under the old acervuli. The pycnidia then overwinter in the lesions of infected tissue and burst in the spring, releasing conidia to be dispersed by water and effectively completing the disease cycle. ''Diplocarpon rosae'' also has a sexual stage, although this is rarely observed in North America due to unfavorable environmental conditions. In this stage, the sexual spores (ascospores) are formed in the apothecium. If the weather conditions are favorable for the formation of ascocarps, the apothecia that contain asci can be observed in the spring. However, this rarely occurs, and the fruiting bodies are typically filled with conidia that enable the asexual life cycle of the pathogen to occur.


Environment

''Diplocarpon rosae'' typically favor environments with a warm and wet climate. Conidiospores involved with infection are only dispersed via water, making the disease most active in the late spring and early fall seasons, or other periods that experience similar climate conditions. The development of the Black Spot disease itself is ideal at temperatures ranging from .Drewes-Alvarez R.(2003) Disease/black spot. In: ''Encyclopedia of Rose Science'' pp 148–153.Eds A.V.Roberts, T.Debener and S.Gudin. Amsterdam, the Netherlands: Elsevier Academic Press. It is important to note that no infection will develop if the leaf surfaces dry out within 7 hours of the initial conidial germination. Similarly, temperatures above also halt the spread of disease.


Treatment

Removing infected leaves from the plant and fallen leaves from the ground will slow the spread of the infection, as does avoiding wetting the leaves of plants during watering. An infected plant can be removed from the area, which will slow the spread of infection to other plants, but this often is not desirable. Fungicides, such as
mancozeb Mancozeb is a dithiocarbamate non-systemic agricultural fungicide with multi-site, protective action on contact. It is a combination of two other dithiocarbamates: maneb and zineb. The mixture controls many fungal diseases in a wide range of f ...
, chlorothalonil, flutriafol, penconazole, or a copper-based product, applied upon new leaf emergence or first appearance of black spot, can be used to control the disease. If a more natural and nontoxic approach is desired, diluted
neem oil Neem oil, also known as margosa oil, is a vegetable oil pressed from the fruits and seeds of the neem (''Azadirachta indica''), a tree which is indigenous to the Indian subcontinent and has been introduced to many other areas in the tropics. It i ...
is effective both against black spot and as an insecticide against
aphids Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A t ...
, or cows milk diluted 1:3 with water is effective if sprayed on to the leaves. It is usually necessary to repeat the spraying at seven- to 10-day intervals throughout the warmest part of the growing season, as the fungus is most active at temperatures from .


Importance

Black Spot of rose is the single most impactful disease of roses globally. Every year around 8 billion flowering stems, 80 million potted plants and 220 million garden rose plants are sold commercially. All species of roses (''Hulthemia, Hesperrhodos, Platyrhodon'' and Rosa) are affected by black spot disease. The disease is found everywhere roses are planted, typically in epidemic proportions. The water-borne dispersal methods allow it to infect a plethora of plants every growing season and increase the overall incidence of disease. Although ''Diplocarpon'' ''rosae'' does not kill the rose itself, it is known to completely defoliate the leaves of the rose plant. This is a huge issue when dealing with such an aesthetically commercialized crop such as the rose. Additionally, the weakened rose plant will become more susceptible to other pathogens and disease following infection.Gachomo E.W. (2005) ''Studies of the Life Cycle of'' Diplocarpon rosae Wolf on Roses and the Effectiveness of Fungicides on Pathogenesis. Goettingen, Germany: Cuvillier Verlag.


References


External links


Black spot of roses at the University of Maine


{{Taxonbar, from=Q1608118 Dermateaceae Fungi described in 1912 Fungal plant pathogens and diseases Rose diseases