Black Caracara
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The black caracara (''Daptrius ater'') is a species of
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the family
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
found in Amazonian and
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the
Republic of Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. Borders of Suriname, It is bordered by the Atlanti ...
or ''juápipi '' by the Emberá of
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
and
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
. Both these names refer to multiple bird species within Falconidae. German-Brazilian ornithologist
Helmut Sick Helmut Sick (10 January 1910 – 5 March 1991) was a German-Brazilian ornithologist. Sick was born in Leipzig, Germany. He emigrated to Brazil in 1939. A prominent ornithologist in Brazil, Sick published more than 200 papers, including his most ...
also referred to this species as ''gavião-de-anta'', literally translating to "tapir-hawk". Since migration has not been observed, they are considered
sedentary Sedentary lifestyle is a lifestyle type, in which one is physically inactive and does little or no physical movement and or exercise. A person living a sedentary lifestyle is often sitting or lying down while engaged in an activity like soci ...
; remaining in the tropics year round. Its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Daptrius'' is often considered
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
today, though some also include its close relative, the
red-throated caracara The red-throated caracara (''Ibycter americanus'') is a social species of bird of prey in the family Falconidae. It is placed in the monotypic genus ''Ibycter'', or sometimes united in ''Daptrius'' with the black caracara. Unique among caracaras ...
otherwise separated in ''Ibycter''. ''Daptrius ater'' are considered to be a widespread raptor, often spotted in groups of 2-5 individuals in tall trees. Nests built from sticks containing 2-3 spotted, brown eggs have been observed high up in trees, however little else is known about their breeding habits and reproduction. Typical of caracaras, this species is an omnivore as well as an opportunistic feeder, known to be both a predator,
scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...
and forager.


Taxonomy

The species ''Daptrius ater'' was first described by Louis Jean Pierre Vieillot in 1816. It previously shared its genus with the red-throated caracara, which is now known as '' Ibycter americanus'' in the monotypic genus ''Ibycter''. The inclusion of ''I. americanus'' in the genus ''Daptrius'' was made by
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
Herbert Friedmann Herbert Friedmann (April 22, 1900 – May 14, 1987) was an American ornithologist. He worked at the Smithsonian Institution for more than 30 years. In 1929 he became a fellow of the American Ornithologists' Union (AOU) and served as the Presiden ...
in 1950, and backed up by
Dean Amadon Dean Arthur Amadon (June 5, 1912 – January 12, 2003) was an American ornithologist and an authority on birds of prey. Amadon was born in Milwaukee, Wisconsin to Arthur and Mary Amadon. He received a BS from Hobart College in 1934 and a Ph.D. ...
in 1968. However, a recent
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
and nuclear sequence data study focusing on caracaras and allies (subfamily ''Polyborinae'') determined that these two species not only differ in genus, but are also arranged in
polyphyly A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
, meaning that they do not share a common most recent ancestor. Additionally, it was concluded that ''D. aters closest relative is the
yellow-headed caracara The yellow-headed caracara (''Daptrius chimachima'') is a bird of prey in the family Falconidae. It is found in tropical and subtropical South America and the southern portion of Central America. Unlike the falcons in the same family, the caraca ...
(''Milvago chimachima''), existing on the
phylogenetic tree A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
as a sister group.


Description

Adult black caracara are a glossy black except for the distinctive white band on the base of the tail, and yellow to orange-red feet and face. The adults appear similar to those of their closest relative, the
red-throated caracara The red-throated caracara (''Ibycter americanus'') is a social species of bird of prey in the family Falconidae. It is placed in the monotypic genus ''Ibycter'', or sometimes united in ''Daptrius'' with the black caracara. Unique among caracaras ...
, however they have distinctly long and narrow wings and tail, as well as a black beak. Additionally, the red-throated caracara can be distinguished from the black caracara by their red throats. The average adult length is 41-47cm. The females of this species average 350-440g and are typically larger than the males that have an average weight of 330g. Juveniles can be identified by their dull black
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
, pale yellow face and the 3-4 black bars found on the rectrices. When observing flight from a distance, it is notable that ''Daptrius ater'' rarely soars, but instead can be seen continuously flapping.


Habitat and distribution

It is found in
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
,
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
,
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
,
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
,
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
,
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
, and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. Its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are
subtropical or tropical moist lowland forest Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature. Description TSMF is generally found in large, discont ...
and heavily degraded former forest. Common habitats include
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
and wooded
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
that are situated between altitudes of 0 to 900m. Black caracara can also be found in
mangroves A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in several ...
and disturbed forested habitats along water.


Conservation status

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
estimates a fluctuating population of 1000-10000 individuals, with an adult population of 670-6700. Population declines have been documented over the past 10 years, however it is not presently considered critical. The designation of least concern is attributed to their large range, ability to survive in fragmented forest, and their diverse diet.


Behavior

The black caracara are most often seen in pairs or family groups of 3-4, but can be spotted alone. They have been observed flying in straight patterns with continuous flapping, walking along rivers, and perching in tall trees. Other common sightings have famously associated them with
tapir Tapirs ( ) are large, herbivorous mammals belonging to the family Tapiridae. They are similar in shape to a pig, with a short, prehensile nose trunk. Tapirs inhabit jungle and forest regions of South and Central America, with one species inhabit ...
and
capybara The capybaraAlso called capivara (in Brazil), capiguara (in Bolivia), chigüire, chigüiro, or fercho (in Colombia and Venezuela), carpincho (in Argentina, Paraguay and Uruguay) and ronsoco (in Peru). or greater capybara (''Hydrochoerus hydro ...
, as they have been observed picking ectoparasites from the fur. This interaction can be considered mutualistic as tapirs notably solicit nearby black caracara using a call, then lay still to facilitate tick removal. They are also known to scavenge around human settlements and exhibit the ability to fish along rivers.


Diet

''D. ater'' are known to eat almost anything from vegetation to
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
to live prey; therefore, are best described as an opportunistic feeder. More specifically, their diet consists of nestlings and fledglings of other bird species, smaller birds such as flycatchers and pigeons, small mammals, carrion, frogs, reptiles, invertebrates, small fish, palm nuts and other fruit. With this diverse list that makes up their diet, they have developed various foraging and hunting strategies. These include directly attacking the nests of other birds, searching the canopy foliage with their beaks for insects, and exhibiting comfort around humans when scavenging in camps.


Fishing

The diverse list of feeding strategies includes foraging small fish within their riverine habitats. ''D. ater'' make use of exposed rocks and emergent vegetation in and along rivers, walking slowly on these surfaces while peering into the water for small fish such as species of
Characidae Characidae, the characids or characins is a family of freshwater subtropical and tropical fish, belonging to the order Characiformes. The name "characins" is the historical one, but scientists today tend to prefer "characids" to reflect their st ...
. Once the prey is spotted, they are caught using either the bill or
talons A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...
then carried away from the river for consumption. Black caracara mostly fish in areas of fast moving rivers where migratory fish are forced to bottleneck and become trapped among plants or within shallow pools, reflecting their opportunistic nature.


Vocalization

This species can also be identified from other species by their harsh, piercing, single note ''kraaaa'' calls that usually occur during flight. This call is usually repeated several times, often ending in a decrescendo. They do not exhibit diverse vocalizations, restricted to variations in length and volume of their distinctive shriek.


Reproduction

There is only one observational record of a black caracara nest in Brazil being built from twigs 60–70 cm in length, 25m high in a tree. Little else is known about their reproductive behaviors.


References

{{Taxonbar, from=Q520013
black caracara The black caracara (''Daptrius ater'') is a species of bird of prey in the family Falconidae found in Amazonian and French Guiana lowlands, commonly along rivers. They are locally referred to as ''Ger' futu busikaka'' in the Republic of Suriname ...
Birds of the Amazon Basin Birds of Colombia Birds described in 1816 Taxa named by Louis Jean Pierre Vieillot Taxonomy articles created by Polbot