Black And Yellow Mud Dauber
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''Sceliphron caementarium'', also known as the yellow-legged mud-dauber wasp, black-and-yellow mud dauber (within the US), or black-waisted mud-dauber (outside of the US), is a species of
sphecid wasp The Sphecidae are a cosmopolitan family of wasps of the suborder Apocrita that includes sand wasps, mud daubers, and other thread-waisted wasps. The name Sphecidae was formerly given to a much larger grouping of wasps. This was found to be p ...
. There are some 30 other species of ''Sceliphron'' that occur throughout the world, though in appearance and habits they are quite similar to ''S. caementarium''.


Etymology

The Latin species name ''caementarius'' means ''mason'' or ''builder of walls''.


Distribution and habitat

''S. caementarium'' is widespread in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, the
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,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
and the
West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...
, and has been introduced to many
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(including
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
, and
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
),
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
and
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, where it has become established in some countries of the
Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
(
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
and
Corsica Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
,
Malta Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...
, the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
, and
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
) and
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
, and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
. This species is found in a wide variety of habitats, such as rock ledges, man-made structures, puddles and other water edges,
cypress dome A cypress dome is a type of freshwater forested wetland, or a swamp, found in the southeastern part of the United States. They are dominated by the ''Taxodium spp''., either the bald cypress (''Taxodium distichum''), or pond cypress (''Taxodium asc ...
s, in long leaf pines (''
Pinus palustris The longleaf pine (''Pinus palustris'') is a pine species native to the Southeastern United States, found along the coastal plain from East Texas to southern Virginia, extending into northern and central Florida. In this area it is also known as ...
''), and in
turkey oak Turkey oak is a common name for several species of oaks and may refer to: *''Quercus cerris'', native to southeastern Europe and Asia Minor *''Quercus laevis ''Quercus laevis'', the turkey oak, is a member of the red oak group of oaks. It is nat ...
s.


Description

''Sceliphron caementarium'' can reach a length of . Their petiole is generally black and is about half the length of the entire abdomen, however the population in the
desert southwest The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural list of regions of the United States, region of the United States that generally includes Arizona, New Mexico, and adjacen ...
often has a yellow petiole. The thorax shows various yellow markings, while the abdomen is normally black, with yellow
propodeum The propodeum or propodium is the first abdominal segment in Apocrita Hymenoptera (wasps, bees and ants). It is fused with the thorax to form the mesosoma. It is a single large sclerite, not subdivided, and bears a pair of spiracles. It is strongl ...
(typical of females). The eyes are black, the antennae are black, and the legs are yellow with black
trochanter A trochanter is a tubercle of the femur near its joint with the hip bone. In humans and most mammals, the trochanters serve as important muscle attachment sites. Humans are known to have three trochanters, though the anatomic "normal" includes ...
s and femurs. Within the United States, it is the only species with yellow-marked legs. The wings are a tawny color.Bugguide
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Biology

The black and yellow mud daubers are solitary
parasitoid wasp Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causin ...
s that build nests out of mud. These sphecid wasps collect mud balls at puddle and pool edges for constructing nests. Frequently, nests are built in shaded areas inside formations that are sheltered from the weather or from other environmental elements. These sites may be naturally-occurring, or man-made structures. Some examples are: under and inside various types of bridges, barns, garages, open-air porches, or under housing eaves. The nests comprise up to 25 vertically arranged, individual cylindrical cells. After initial creation and covering of the clutch, this sphecid wasp uses more mud as a means covering and protecting the whole cluster of cells, thereby forming a smooth appearance, and a uniform nest. The entire nest may attain an area equal to, or larger than, the size of an average human fist. After building a cell of the nest, the female wasp captures several
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s. The captured prey are stung and paralyzed before being placed in the nest (usually 6-15 per cell), and then a single egg is deposited on the prey within each cell. The wasp then seals the cell with a thick mud plug. After finishing a series of cells, she leaves and does not return. While consuming the prey and increasing in size, the larva molts several times, until it molts into a
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
. Once the pupa has developed into an adult wasp, the adult emerges from its pupal case and breaks out of its mud chamber. Adults can be seen in mid-summer feeding on nectar at flowers, especially Queen Anne's lace (''
Daucus carota ''Daucus carota'', whose common names include wild carrot, European wild carrot, bird's nest, bishop's lace, and Queen Anne's lace (North America), is a flowering plant in the family Apiaceae. It is native to temperate regions of the Old Wor ...
''), parsnips and water parsnips (''
Sium suave ''Sium suave'', the water parsnip or hemlock waterparsnip, is a perennial wildflower in the family Apiaceae. It is native to many areas of both Asia and North America. The common name water parsnip is due to its similarity to parsnip (''Pastinaca ...
'', ''
Sium latifolium ''Sium latifolium'' is a species of flowering plant in the family Apiaceae known by the common names great water-parsnip, greater water-parsnip, and wideleaf waterparsnip. It is native to much of Europe, Kazakhstan, and Siberia. This plant grows ...
'', ''
Berula erecta ''Berula erecta'', known as lesser water-parsnip or cutleaf waterparsnip or narrow-leaved water-parsnip, is a member of the carrot family. Growing to around tall, it is found in or by water. It is widespread across much of Europe, Asia, Austral ...
''). They have a low reproductive rate. Stings are rare due to their solitary and usually nonaggressive nature; however, nests are aggressively defended. A common species of
cuckoo wasp Commonly known as cuckoo wasps or emerald wasps, the hymenopteran family Chrysididae is a very large cosmopolitan group (over 3000 described species) of parasitoid or kleptoparasitic wasps, often highly sculptured, with brilliant metallic colors ...
, '' Chrysis angolensis'', is frequently a
cleptoparasite Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when foo ...
in ''
Sceliphron ''Sceliphron'', also known as black mud daubers or black mud-dauber wasps, is a genus of Hymenoptera of the Sphecidae family of wasps. They are solitary mud daubers and build nests made of mud. Nests are frequently constructed in shaded niches, o ...
'' nests, and is only one of many different insects that parasitize these
mud dauber Mud dauber (or "mud wasp" or "dirt dauber") is a name commonly applied to a number of wasps from either the family Sphecidae or Crabronidae which build their nests from mud; this excludes members of the family Vespidae (especially the subfamily Eu ...
s.


Venom

Although they are common components of
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
s,
serotonin Serotonin () or 5-hydroxytryptamine (5-HT) is a monoamine neurotransmitter. Its biological function is complex and multifaceted, modulating mood, cognition, reward, learning, memory, and numerous physiological processes such as vomiting and vas ...
,
histamine Histamine is an organic nitrogenous compound involved in local immune responses, as well as regulating physiological functions in the gut and acting as a neurotransmitter for the brain, spinal cord, and uterus. Since histamine was discovered in ...
,
acetylcholine Acetylcholine (ACh) is an organic chemical that functions in the brain and body of many types of animals (including humans) as a neurotransmitter. Its name is derived from its chemical structure: it is an ester of acetic acid and choline. Part ...
, and
kinin A kinin is any of various structurally related polypeptides, such as bradykinin and kallidin. They are members of the autacoid family. Kinins are peptides that are cleaved from kininogens by the process of kallikreins. Kallikreins activate kinins wh ...
s are absent from ''S. caementarium'' venom.


Mentions in popular media

In 1996,
Birgenair Flight 301 Birgenair Flight 301 was a flight chartered by Turkish-managed Birgenair partner Alas Nacionales from Puerto Plata in the Dominican Republic to Frankfurt, Germany, via Gander, Canada, and Berlin, Germany. On February 6, 1996, the 757-200 oper ...
crashed near Puerto Rico. The most probable cause of this
Boeing 757 The Boeing 757 is an American narrow-body airliner designed and built by Boeing Commercial Airplanes. The then-named 7N7, a twinjet successor for the 727 (a trijet), received its first orders in August 1978. The prototype completed its mai ...
crash was a blockage in a
Pitot tube A pitot ( ) tube (pitot probe) measures fluid flow velocity. It was invented by a French engineer, Henri Pitot, in the early 18th century, and was modified to its modern form in the mid-19th century by a French scientist, Henry Darcy. It is ...
(device for air speed measurement) by a mud dauber's nest.


Gallery

File:Sceliphron-mud-dauber.jpg, ''S. caementarium'' collecting mud File:20100710.MudDauber-SceliphronCaementarium.Hannibal.jpg, ''S. caementarium'' with a load of mud, just before takeoff File:Black and Yellow Mud Dauber (Sphecidae, Sceliphron caementarium (Drury)) (36164276721).jpg, ''S. caementarium'' with a partially yellow petiole from Austin, TX File:Sphecidae - Sceliphron caementarium - Nest.JPG, A nest containing the cells covered with mud File:Contenuto di un nido di Sceliphron Caementarium (Vespa muratore).jpg, Paralyzed spiders used as larval provisions collected from a nest


References


Further reading

*Ascher J.S., Pickering J. (2016) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila) *Bohart, R.M., and A.S. Menke. 1976. Sphecid Wasps of the World: a Generic Revision. University of California Press. *Crawford, R.L. 1987. Spider prey of the mud-dauber, Sceliphron caementarium (Sphecidae), in Washington. Proceedings of the Washington State Entomological Society, 48: 797-800. *Evans, H.E., and M.J.West-Eberhard. 1970. The Wasps. University of Michigan Press. *Howard, L.O. 1903. The Insect Book. Doubleday. *Kevin M. O'Neill. 2000 - Solitary Wasps: Behavior and Natural History (Cornell Series in Arthropod Biology) - Comstock Publishing *Rau, Phil. 1916. The sleep of insects. Annals of the Entomological Society of America, 9: 227-275. *Roques A., Kenis M., Lees D., Lopez-Vaamonde C., Rabitsch W., Rasplus J.-Y., Roy D. - Alien terrestrial arthropods of Europe - BioRisk 4 Special Issue *Ross H. Arnett - American Insects: A Handbook of the Insects of America North of Mexico - CRC Press. *Ross, K., and R. Matthews. 1991. The Social Biology of Wasps. Chapter 17, Evolution of Social Behavior in Sphecid Wasps, pp. 570–602.


External links

* {{DEFAULTSORT:Black And Yellow Mud Dauber Sphecidae Biological pest control wasps Insects described in 1773 Hymenoptera of Europe