Black-throated Gray Warbler
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The black-throated gray warbler or black-throated grey warbler (''Setophaga nigrescens'') is a passerine bird of the New World warbler family Parulidae. It is long and has gray and white plumage with black markings. The male has the bold black throat of its name, and black stripes on its head, as well as black streaks on its flanks; the female is a paler version of the male, with a white throat and less distinct black markings on the flanks and wings. It breeds in western North America from British Columbia to New Mexico, and winters in Mexico and the southwestern United States. The habitats it prefers are coniferous and mixed forests and scrubland, especially those with pinyon pines,
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
s, sagebrush, and oaks. Its nest is an open cup of plant fibers lined with feathers, built a few metres from the ground in the branches of a tree or shrub. Three to five eggs are laid, and young are fed by both parents. Common in its breeding range, it does not seem to be seriously threatened by human activities, unlike many migratory warblers.


Taxonomy

The black-throated gray warbler was first described by John Kirk Townsend from a specimen collected near today's Portland, Oregon. It was known to the Chinook inhabitants of the northwest coast, who called it ''Ah Kah a qual''. Townsend described the species as ''Sylvia nigrescens'', placing it with the other New World warblers and the unrelated Old World warblers in the genus '' Sylvia''. It is now placed in the genus '' Setophaga'' along with about thirty other species, after having been classified in ''Dendroica'' for some time. Within its genus, it is part of a group with black throats and yellow face markings that includes the
hermit warbler The hermit warbler (''Setophaga occidentalis'') is a small perching bird. It is a species of New World warbler or wood-warbler. They are a migratory bird, the breeding range spanning the majority of the west coast of the United States. Their wint ...
and
Townsend's warbler Townsend's warbler (''Setophaga townsendi'') is a small songbird of the New World warbler family. Taxonomy Townsend's warbler was species description, formally described in 1837 by the American naturalist John Kirk Townsend under the binomial nam ...
. It is usually considered an early offshoot among this group of species, but genetic studies suggest a close relation to
Grace's warbler Grace's warbler (''Setophaga graciae'') is a small New World warbler that specializes in pine woods. Taxonomy Grace's warbler was discovered by Elliott Coues in the Rocky Mountains in 1864. He requested that the new species be named after his 18 ...
. Of these relatives, the range of the black-throated gray warbler overlaps with those of Townsend's warbler and the hermit warbler, but they occur in different habitats. While Townsend's and hermit warblers commonly hybridize with each other, records of either species hybridizing with the black-throated gray warbler are uncommon. There are two
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, which are highly similar and of dubious validity. The
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''S. n. nigrescens'' occurs on the Pacific coastal region from British Columbia to northern California, while ''S. n. halseii'', described by Giraud in 1841, occupies the inland parts of its range. ''S. n. halseii'' birds are slightly larger and more gray in their upperparts.


Description

The black-throated gray warbler has mostly black, gray, and white plumage, which is soft, lacking gloss. With its striping and the small yellow spot between its eye and bill, it is a distinctive bird. The sexes differ slightly, both having gray upperparts with black streaks, and white underparts with black streaks on the flanks. The adult male is striped with black on the crown, throat, and below the eye, and has white around its chin and above its eye. The adult female has more dingy plumage on its head, with a white throat and dark gray cheeks. The most similar birds to the black-throated gray warbler are the black-and-white and blackpoll warblers, which although marked in black have entirely different plumage patterns. It is typically long, weighing . Wing lengths are , tail lengths , bill lengths , and tarsus lengths , with females slightly smaller than males. Wingspan ranges from 7.5-7.8 in (19-19.7 cm). This bird gives a sharp ''tup'' or ''thick'' call, like that of
Townsend's warbler Townsend's warbler (''Setophaga townsendi'') is a small songbird of the New World warbler family. Taxonomy Townsend's warbler was species description, formally described in 1837 by the American naturalist John Kirk Townsend under the binomial nam ...
but flat and unmusical, as well as a high ''see''
flight call Flight calls are vocalisations made by bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a fou ...
. The male's song is a series of buzzy notes, with the earlier notes doubled and the next to last note high. This song has three variations, including a quiet "soft song" given by the males while following females gathering material for a nest.


Distribution and habitat

The black-throated gray warbler breeds in western North America, its range extending from southwestern British Columbia along the Pacific coast to northern Baja California, and east to New Mexico and southern Montana. It winters mostly in Mexico, from southern Baja California to Oaxaca state. It has spread into parts of Wyoming and Montana only recently, as '' Juniperus osteosperma'' has expanded its range due to a warming climate. Vagrants have been recorded across eastern North America and in Cuba. The black-throated gray warbler breeds in open
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
and mixed forest with a brushy understory, in dry open oak forests, and in
chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...
and other scrubland. It is particularly associated with pinyon pines,
juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' () of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere, from the Arcti ...
s, and oaks. It migrates to the south late in the fall, returning north in mid-spring. While migrating, it forages in any woodland or scrub it passes through. In its wintering grounds, it occurs in dry woodland and tall scrub. Though its status is not well known, it does not appear to be seriously threatened by habitat destruction or other human activities, unlike many migratory warblers. It is a fairly common bird, among the most common in some localities. Because it is common and not in rapid decline, it is assessed as Least Concern on the IUCN Red List.


Behavior

The black-throated gray warbler is usually approachable, and can easily be observed while foraging. Despite this, it is poorly known, especially in its breeding behavior, of which '' Birds of North America'' said "almost no information is available". It forages, often in flocks with other species. It feeds on insects
gleaned Gleaning is the act of collecting leftover crops from farmers' fields after they have been commercially harvested or on fields where it is not economically profitable to harvest. It is a practice described in the Hebrew Bible that became a legall ...
on low branches, especially
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s. The nest is usually placed on a horizontal tree branch or in a shrub, a few metres above the ground. The nest is an open cup constructed of grass stalks and other fibres, and lined with feathers and hair. The female lays three to five pinkish eggs with brown dots from May to July. Incubation and fledging periods are unknown. This species has been recorded giving a distraction display, pretending to be injured to distract predators from its nest. Both parents feed the young, though the female may do so more frequently.


References


Works cited

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External links


Black-throated Gray Warbler
at All About Birds
Black-throated Grey Warbler
at the Internet Bird Collection {{Taxonbar, from=Q3176524 black-throated gray warbler Native birds of Western Canada Native birds of the Western United States black-throated gray warbler black-throated gray warbler