Black-tailed Skimmer
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The black-tailed skimmer (''Orthetrum cancellatum'') is a
dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
belonging to the family
Libellulidae The skimmers or perchers and their relatives form the Libellulidae, the largest dragonfly family in the world. It is sometimes considered to contain the Corduliidae as the subfamily Corduliinae and the Macromiidae as the subfamily Macromiinae. Ev ...
.


Distribution

This species is widespread in Europe and Asia. It is found throughout European continent including the Mediterranean islands but is absent in the north of Britain and the northern half of Fennoscandia. This is one of the most common European species and it is still increasing its range northwards. To the east, the range extends over central Asia to Kashmir, Mongolia to the northern parts of China and Arunachal Pradesh in India. It is present in
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and in part of
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
. The black-tailed skimmer is abundant throughout its range and is one of the most commonly seen dragonflies in Europe. It holds a stable population and has no known major threats. This species has expanded its range, assisted by the creation of
gravel pit A gravel pit is an open-pit mine for the extraction of gravel. Gravel pits often lie in river valleys where the water table is high, so they may naturally fill with water to form ponds or lakes. Old, abandoned gravel pits are normally used eithe ...
s which give it the extensive open unvegetated areas it prefers. It was first recorded in
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
in Essex in 1934. It is decreasing rapidly in the
Maltese Islands Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...
.


Habitat

This dragonfly is found at any open water with bare patches along the shore where the patrolling males frequently rest in the sun. It also inhabits near slow-flowing waters. It favors lakes, slow rivers, ponds and sometimes marshy area, without dense riparian vegetation. Females are less bold and not encountered as regularly. Adults prefer to perch on bare ground and rocks.


Description

It is a fairly large dragonfly (the length of 47–53 mm, 29–35 mm abdomen, rear wing 35–41 mm.) with relatively broad, flattened abdomen, but not as broad as to chaser species. With age, adult males develop extensive blue pruinescence on their abdomen, offset by yellow lateral patches. The middle lobe of the
pronotum The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum (dorsal), the prosternum (ventral), and the propleuron (lateral) on ea ...
is large and notched in the middle. The chest is yellow or yellowish-brown. The base of the hind wings do not show a dark opaque spots. The
pterostigma The pterostigma (plural: pterostigmata) is a group of specialized cells in the outer insect wing, wings of insects, which are often thickened or coloured, and thus stand out from other cells. It is particularly noticeable in dragonfly, dragonfli ...
is dark brown or black. On the front wings pterostigma 2–3 mm long. Anal appendages are black. The females and immature males are a deep yellow color, with wavy black lines dorsally on their abdomen. Males and females have the
costal vein Costal may refer to: * an adjective related to the rib () in anatomy ** Costal cartilage, a type of cartilage forming bars which serve to prolong the ribs forward ** Costal margin, the medial margin formed by the false ribs ** Costal surface (disa ...
(the leading edge of the wing) yellow or black. The species is similar to its much more localized congener the '' Orthetrum albistylum'', but readily identifiable in the field. The males develop a blue
pruinescence Pruinescence , or pruinosity, is a "frosted" or dusty-looking coating on top of a surface. It may also be called a pruina (plural: ''pruinae''), from the Latin word for hoarfrost. The adjectival form is pruinose . Entomology In insects, a "blo ...
on the abdomen darkening to the rear with S8-10 becoming black. Its eyes are very dark green. They fly swift and low, skimming the water surface. Females retain their color and markings though they become quite grayish brown with age. This species could be confused with Keeled skimmer or Scarce chaser.


Adults

♂: Adult males have a bluish or blue-gray
patina Patina ( or ) is a thin layer that variously forms on the surface of copper, brass, bronze and similar metals and metal alloys (tarnish produced by oxidation or other chemical processes) or certain stones and wooden furniture (sheen produced b ...
on the body, gradually darkening with age. They have a blue
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
with a black tip and transparent wings. The young have a grid pattern on a yellow-brownish abdomen. ♀: Adult females have a yellow-brownish abdomen with a lattice pattern (wide longitudinal dark brown stripes and bright crescent-shaped spots) and transparent wings. 10
tergite A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'mar ...
of abdomen is black. Even the abdomen of the immature males looks that way. File:Libellulidae - Orthetrum cancellatum (immature male).JPG, Immature male File:Black-tailed skimmer dragonfly (Orthetrum cancellatum) adult male.jpg, Mature male File:Orthetrum cancellatum top.jpg, Immature female File:Black-tailed skimmer dragonfly (Orthetrum cancellatum) mature female.jpg, Mature female Adults can reach a length of about 47–53 mm. Average wingspan is about 77 mm, while hindwings are 35–40 mm long. The eyes are dorsally narrowly apposed to
dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
broadly contiguous. They are brown (in the female), or blue (in the male). Legs are black and brown (the femoral bases brown). Thoracic antehumeral stripes are absent. The wings spread more or less horizontally in repose. They are dissimilar in shape and venation, sessile, unpatterned and clear. The inner wing
venation Venation may refer to: * Venation (botany), the arrangement of veins in leaves * Wing venation, the arrangement of veins in insect wings See also * * Vernation Vernation (from ''vernal'' meaning ''spring'', since that is when leaves spring forth ...
is blackish. Discoidal cell is divided longitudinally into a conspicuous triangle and supra-triangle. There are antenodal veins in the forewings 10–13 (fewer in the hindwings).
Pterostigma The pterostigma (plural: pterostigmata) is a group of specialized cells in the outer insect wing, wings of insects, which are often thickened or coloured, and thus stand out from other cells. It is particularly noticeable in dragonfly, dragonfli ...
is well over twice but no more than five times as long as wide. Abdomen is lanceolate and 30–35 mm long; predominantly blue (in the male), or brown (in the female), or grey (sometimes, in old females). It is rather plain (in the male, but often with orange lateral markings, and darkened over the terminal segments), or predominantly longitudinally lined (the female with a dark medio-lateral band on either side along each segment); without mid-dorsal spots. The male abdomen is without auricles on segment 2; with a single inferior anal appendage.


Nymphs

The
nymphs A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are typ ...
are stout, body is expanded in the middle. When mature, they are 23–25.5 mm long. The postocular lobes is curving sharply to the back of the head from some distance behind the eyes. The antennae is 7 segmented. The mask have the prementum hollowed dorsally. The prementum bears three (rarely two or four) long setae inserted near each lateral margin, and two medial fields of seven to eight short spiniform setae, the latter flanked by four setae of medium length. The body of the labial palps bears 6 or 7 major setae. Distal margins of the labial palps are weakly crenate. Legs are longer than the abdomen. The abdomen terminates in five short spine-like appendages. The cerci are no more than half the length of the paraprocts. The abdomen has mid-dorsal spines. The mid-dorsal abdominal spines are prominent on segments 3 to 6. The abdomen is without a mid-dorsal spine on segment 8; gizzard with 4–8 folds.


Biology and behavior

The main flight period is June and July. Males characteristically perch horizontally on exposed surfaces. They fly swift and low, skimming the water surface, while defending their territories. Mating can occur in flight or on land. Females
oviposit The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...
alone but with male in attendance. Eggs are laid in flight by dipping abdomen onto water surface. They hatch after five or six weeks and the larvae live partially hidden by bottom debris. They emerge after two or three years. Larvae prefer bottom areas rich in vegetation and decaying plant material.


References


Sources

* * Klaas-Douwe B Dijkstra, Richard Lewington, 2006: ''Field Guide to the Dragonflies of Britain and Europe.'' British Wildlife Publishing, Gillingham. * A. Sciberras, 2008: ''A Contribution To The Knowledge Of Odonata In The Maltese Islands.'' The Central Mediterranean Naturalist 4(4): 275-288.


External links


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{{DEFAULTSORT:Black-Tailed Skimmer Libellulidae Dragonflies of Europe Insects described in 1758 Taxa named by Carl Linnaeus