Black-headed Grosbeak
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The black-headed grosbeak (''Pheucticus melanocephalus'') is a medium-sized,
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
-eating
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the family
Cardinalidae Cardinalidae (often referred to as the "cardinal-grosbeaks" or simply the "cardinals") is a family of New World-endemic passerine birds that consists of cardinals, grosbeaks, and buntings. It also includes several birds such as the tanager-like ...
. It is sometimes considered conspecific with the
rose-breasted grosbeak The rose-breasted grosbeak (''Pheucticus ludovicianus''), colloquially called "cut-throat" due to its coloration, is a large, seed-eating grosbeak in the cardinal family (Cardinalidae). It is primarily a foliage gleaner. Males have black heads, w ...
(''P. ludovicianus'') with which it hybridizes on the American
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, an ...
. The long, black-headed grosbeak is a
migratory bird Bird migration is the regular seasonal movement, often north and south along a flyway, between Breeding in the wild, breeding and wintering grounds. Many species of bird migrate. Animal migration, Migration carries high costs in predation a ...
, with nesting grounds from southwestern
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
, through the western half of the United States, into central
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. It occurs as a
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
further south in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
.


Description

Measurements: * Length: * Weight: * Wingspan: The black-headed grosbeak is similar in size to a
common starling The common starling or European starling (''Sturnus vulgaris''), also known simply as the starling in Great Britain and Ireland, is a medium-sized passerine bird in the starling family, Sturnidae. It is about long and has glossy black plumage ...
. As per its name, the male has a black head, and black wings and tail with prominent white patches. Its breast is dark to tawny orange in color, and its belly is yellow. The female has a brown head, neck, and back with sparrow-like black streaks. She also has white streaks down the middle of her head, over her eyes, and on her cheeks. Her breast is white, and wings and tail are grayish-brown with two white wing bars and yellowish wing edges.


Habitat

The black-headed grosbeak prefers to live in deciduous and mixed wooded areas. It likes to be in areas with large trees and thick bushes, such as patches of broadleaved trees and shrubs within conifer forests, including streamside corridors, river bottoms, lakeshores, wetlands, and suburban areas.


Nesting

Females build nests among the dense foliage on an outer branch of tall broadleaved trees or shrubs, above ground. They occasionally build in dense shrubs such as
blackberry The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of ...
. The nest is in the shape of an open saucer, made of fine grass, rootlets, twigs, bark, and conifer needles. It is often lined with rootlets, hair, and fine plant material. The female lays two to five pale green, blue, or gray eggs that are spotted with reddish and dark brown. The eggs are incubated by the male and female for 12–14 days. After the eggs have hatched, the fledglings leave the nest in about 11 or 12 days, but they are unable to fly for another two weeks. The young are fed by both adults. The black-headed grosbeak's monogamy is under study, but pair bonds generally last for only one breeding season. They typically have one brood per season, though two broods have been documented in foothills of the
Sacramento Valley , photo =Sacramento Riverfront.jpg , photo_caption= Sacramento , map_image=Map california central valley.jpg , map_caption= The Central Valley of California , location = California, United States , coordinates = , boundaries = Sierra Nevada (ea ...
in California.


Voice

The grosbeak's song is a rich warble that is similar to that of an
American robin The American robin (''Turdus migratorius'') is a migratory bird of the true thrush genus and Turdidae, the wider thrush family. It is named after the European robin because of its reddish-orange breast, though the two species are not closel ...
, but more fluent, faster, softer, sweeter, and mellow with rising and falling passages that make the song much longer than the robin's. The note is a sharp ''ik'' or ''eek''. Both the male and female sing, but have different songs.


Diet

The black-headed grosbeak eats pine and other seeds, berries, insects and spiders, and fruit. During the summer, it mostly eats
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s, snails, and
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s. It is one of the few birds, along with the
black-backed oriole The black-backed oriole (''Icterus abeillei'') is a species of bird in the family Icteridae. It is endemic to Mexico. Its natural habitats are subtropical or tropical moist lowland forest and subtropical or tropical moist montane forest, but i ...
, that can safely eat the poisonous
monarch butterfly The monarch butterfly or simply monarch (''Danaus plexippus'') is a milkweed butterfly (subfamily Danainae) in the family Nymphalidae. Other common names, depending on region, include milkweed, common tiger, wanderer, and black-veined brown. It ...
. In their wintering grounds, this grosbeak consumes many
monarch butterflies The monarch butterfly or simply monarch (''Danaus plexippus'') is a milkweed butterfly (subfamily Danainae) in the family Nymphalidae. Other common names, depending on region, include milkweed, common tiger, wanderer, and black-veined brown. It ...
, perhaps over one million per year in the overwintering colonies in Mexico. However, only the black-headed grosbeak is physiologically insensitive to the
cardiac glycoside Cardiac glycosides are a class of organic compounds that increase the output force of the heart and decrease its rate of contractions by inhibiting the cellular sodium-potassium ATPase pump. Their beneficial medical uses are as treatments for co ...
heart poisons in the monarchs, which monarch caterpillars sequester from their milkweed host plants and are retained by adults even after metamorphosis, which protects them from most predaceous birds. The same genetic changes that confer resistance to the poisons in the monarch and its relatives were found to have evolved in the black-headed grosbeak in a case of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
. The cellular target of the heart poisons is the first extracellular loop of the
sodium pump Sodium is a chemical element with the symbol Na (from Latin ''natrium'') and atomic number 11. It is a soft, silvery-white, highly reactive metal. Sodium is an alkali metal, being in group 1 of the periodic table. Its only stable isot ...
, a critical ion pump needed for nerve cells to fire and heart cells to contract. Two of the grosbeak's four copies of the gene encoding this protein have changes that are also found in insects adapted to feeding on plants producing the cardiac glycosides. In one of the copies, it has the same genetic changes at position 111 and 119 as in the common grow and other ''
Euploea ''Euploea'' is a genus of milkweed butterflies. The species are generally dark in coloration, often quite blackish, for which reason they are commonly called crows. As usual for their subfamily, they are poisonous due to feeding on milkweeds and ...
'' butterflies closely related to the monarch, and in another copy, position 111 has the same change as '' Largus'' bugs, which feed on
dogbane Dogbane, dog-bane, dog's bane, and other variations, some of them regional and some transient, are names for certain plants that are reputed to kill or repel dogs; "bane" originally meant "slayer", and was later applied to plants to indicate tha ...
plants that produce cardiac glycosides The mutations at positions 111 and 119 have been shown together, but not alone, to confer resistance to the heart poisons in insects through positive, intramolecualr
epistasis Epistasis is a phenomenon in genetics in which the effect of a gene mutation is dependent on the presence or absence of mutations in one or more other genes, respectively termed modifier genes. In other words, the effect of the mutation is dep ...
. In the breeding season, the black-headed grosbeak comes to bird feeders for sunflower and other types of seed, and fruit, and also joins northern orioles at feeders with grape jelly.


Range and migration

Black-headed grosbeaks range from the Pacific Coast to the middle of the US Great Plains and from southwestern Canada to the mountains of Mexico. US and Canadian birds are highly migratory, wintering in Mexico. In the Great Plains, the range of the black-headed grosbeak and the rose-breasted grosbeak overlap and they have interbred somewhat. After the breeding season, they tend to seek out berry-rich areas. They migrate south early in the fall and return to the north late in the spring and have been known to do so in flocks.


Behavior

Black-headed grosbeaks frequently sing from prominent perches. Both the male and female sing, but have different songs, and both are known to sing from the nest while incubating. When trying to court a female, males fly with their wings and tails spread. They forage in the foliage, on the ground, or in low vegetation and are prominent berry eaters. When feeding the chicks sing to their mother, rustling wings and displaying their yellow under-feathers (see image). File:Blackheaded Grosbeaks.jpg, alt=Photograph of female feeding chick, The birdcall is just delightful. The chick sings to its mother, calling for food; seeds she has cracked open and shelled.


References

* *


Further reading

*


External links


Black-headed grosbeak - ''Pheucticus melanocephalus''
- USGS Patuxent Bird Identification InfoCenter

- Cornell Lab of Ornithology *

* {{Taxonbar, from=Q998547 Grosbeaks Pheucticus Native birds of Western Canada Native birds of the Western United States Birds of Mexico Birds described in 1827