Bemaraha Plateau
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The Bemaraha Formation is a
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
(early
Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratig ...
to early
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
)
geological formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...
of the
Morondava Basin Morondava Basin is one of the major sedimentary basins of Madagascar. It encompasses the large Tsimiroro heavy oil field and giant Bemolanga Bemolanga is a large oil sands deposit in the onshore Morondava Basin of Madagascar. The deposit was ...
of
Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
. The
lime Lime commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a color between yellow and green Lime may also refer to: Botany ...
mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ...
s,
grainstone Under the Dunham classification (Dunham, 1962) system of limestones, a grainstone is defined as a grain-supported carbonate rock that contains less than 1% mud-grade material. This definition has recently been clarified as ''a carbonate-dominated ...
s and
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
s of the formation were deposited in
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') a ...
al and
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock out ...
al
environments Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
. Fossils of groups of invertebrates and
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
and
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
tracks have been found in the formation.Tsiandro tracksite
at
Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...
.org The Sahalaly River tracksite yielded a single trackway belonging to a quadrupedal dinosaur, possibly a sauropod.Wagensommer et al., 2010, p.127 Sauropod tracks are rare, making up as little as 1 percent of the present sample. The best-preserved tracks are assigned to the
ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
'' Kayentapus''.Wagensommer et al., 2012, p.124


Description

The limestones of the Bemaraha Formation have been interpreted as a coastal barrier/lagoon complex of Middle Jurassic age (Bajocian, about 170 Ma). At that time, Madagascar had already separated from Africa, but was still attached to India and Antarctica. Since Antarctica was connected to southern Africa, a
land bridge In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and Colonisation (biology), colonize new lands. A land bridge can be created by marine regre ...
really existed between Madagascar and all major southern hemisphere landmasses. Faunal exchanges were possible and
endemism Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
was probably very low. Madagascar was to become an island only some 80-90 million years later, during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
. During Bajocian times, a narrow seaway stretched between eastern Africa and western Madagascar, much in the same position as today's
Mozambique Channel The Mozambique Channel (french: Canal du Mozambique, mg, Lakandranon'i Mozambika, pt, Canal de Moçambique) is an arm of the Indian Ocean located between the Southeast African countries of Madagascar and Mozambique. The channel is about long ...
. Along the shores of western Madagascar,
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
s separated wide
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') a ...
s from the open sea. The remnants of this reef/lagoon complex is what today, after the coastline shifted several tens of kilometers westward, is seen in the limestones of the Bemaraha Formation. The Bemaraha limestones stretches over an area of roughly ,Wagensommer et al., 2012, p.111 and are best exposed in the area between the towns of
Bekopaka Bekopaka is a rural municipality in western Madagascar. It belongs to the district of Antsalova, which is a part of Melaky Region. The population of the commune was estimated to be approximately 9,000 in 2001 commune census. Bekopaka is served by ...
and
Antsalova Antsalova is a town and commune ( mg, kaominina) in western Madagascar. It belongs to the district of Antsalova, which is a part of Melaky Region. The population of the commune was estimated to be approximately 58,280 in 2018. Antsalova is serve ...
, in the northern half of the
Morondava Basin Morondava Basin is one of the major sedimentary basins of Madagascar. It encompasses the large Tsimiroro heavy oil field and giant Bemolanga Bemolanga is a large oil sands deposit in the onshore Morondava Basin of Madagascar. The deposit was ...
. The outcropping limestones can roughly be divided into a western and an eastern belt with different
facies In geology, a facies ( , ; same pronunciation and spelling in the plural) is a body of rock with specified characteristics, which can be any observable attribute of rocks (such as their overall appearance, composition, or condition of formatio ...
. In the western belt, the formation mainly consists of massive limestones with metre-scale bedding or no recognizable bedding at all.
Oolite Oolite or oölite (''egg stone'') is a sedimentary rock formed from ooids, spherical grains composed of concentric layers. The name derives from the Ancient Greek word for egg (ᾠόν). Strictly, oolites consist of ooids of diameter 0.25–2 ...
s and fossils of reefal organisms (mainly
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
s) are common in places. This region is affected by the spectacular
karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
features known as "
tsingy Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
" and for this reason most of the area is part of the
Tsingy de Bemaraha National Park The Tsingy de Bemaraha National Park is a national park located in Melaky Region, northwest Madagascar. The national park centers on two geological formations: the ''Great Tsingy'' and the ''Little Tsingy''. Together with the adjacent Tsingy de ...
, included in the
UNESCO World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
s list. The facies in the eastern belt is made up mainly of fine-grained and well-bedded limestones with the average thickness of the individual beds being a few tens of centimeters. Beds are often laminated and invertebrate fossils (
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s,
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
s,
brachiopod Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, w ...
s,
echinoid Sea urchins () are spine (zoology), spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard s ...
s and isolated coral fragments) are common in some beds, while others are devoid of fossils.D'Orazi Porchetti et al., 2010, p.8 In the south-western Majunga Basin, interbedded limestones and mudstones (shales and marls) above the Aalenian Sandstone were attributed to a Bajocian carbonate platform formed by the Bemaraha Formation. The carbonate platform at the Manambolo and
Tsiribihina River The Tsiribihina is a river of western Madagascar. The main tributaries are the Mahajilo, Manandaza, Mania, and Sakeny rivers. Its basin has an area of 49,800 km2. 7,025 km2 is in the basin of the Sakeny River, 14,500 km2 in that of the Mahaj ...
gorges, which cut through the Bemaraha plateau, are likewise assigned to the Early Bajocian. At the eastern margin of the Bemaraha Plateau, the carbonate succession directly rests on Isalo sandstones. Farther south at Besabora (along National Road RN35), limestones with the
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular Ectoplasm (cell biology), ectoplasm for catching food and ot ...
'' Mesoendothyra croatica'' and '' Protopeneroplis striata'' support a Middle Jurassic (Bajocian to Early Bathonian) age.Geiger & Schweigert, 2005, p.5


See also

*
List of stratigraphic units with dinosaur tracks Indeterminate or unspecified dinosaur tracks Ornithischians Indeterminate or unspecified ornithischian tracks Saurischians Theropods Sauropodomorphs See also List of dinosaur-bearing rock formations This list of dinosaur-bearing rock f ...
**
List of stratigraphic units with theropod tracks The following tables list the global geological sites where tracks of theropod dinosaurs have been found, together with the proper names of the rock formations (stratigraphic units) that contain them. Non-avian theropods Avialans See also *L ...
**
List of stratigraphic units with sauropodomorph tracks Prosauropods Sauropods See also List of dinosaur-bearing rock formations This list of dinosaur-bearing rock formations is a list of geologic formations in which dinosaur fossils have been documented. Containing body fossils * List of str ...
*
List of fossiliferous stratigraphic units in Madagascar Several fossiliferous stratigraphic units in Madagascar have provided fossils, most notably the Maastrichtian Maevarano Formation and the Jurassic Isalo III Formation. The oldest fossil-bearing formations date to ages before the break-up of Mada ...
*
Geology of Madagascar The geology of Madagascar comprises a variety of rocks of Precambrian age which make up the larger part of the east and centre of the island. They are intruded by basalts and rhyolites of Mesozoic to Cenozoic age. In contrast, the western part of ...
*
Isalo III Formation The Isalo III Formation is a geological formation in Madagascar, off the eastern coast of Africa. It dates back to the Middle Jurassic.Weishampel et al., 2004, pp.542-543 The use of the term "Isalo III" is somewhat controversial as the two prior u ...
*
Kandreho Formation The Kandreho Formation is an Early Jurassic (middle or late Toarcian) geological formation of the Mahajanga Basin of Madagascar. The marly limestones of the formation were deposited in a subtidal lagoonal environment. The formation overlies the ...
*
Etjo Sandstone The Etjo Sandstone is an Early Jurassic geologic Formation (geology), formation in northern Namibia. The formation overlies the Omingonde Formation in the Waterberg Basin and the Doros Formation, Doros and Gai-As Formations in the Huab Basin and h ...


References


Bibliography

* * * * * {{cite LSA , last=Wagensommer , first=A. , last2=Latiano , first2=M. , last3=Nicosia , first3=U. , year=2010 , title=First report of dinosaur footprints from Madagascar: two tracksites from the Middle Jurassic Bemaraha Formation , url=https://www.researchgate.net/publication/233239183_First_Report_of_Dinosaur_Footprints_from_Madagascar_Two_Tracksites_from_the_Middle_Jurassic_Bemaraha_Formation , journal= Ichnos , volume=17 , pages=127–136 , accessdate=2018-08-30 Geologic formations of Madagascar Jurassic System of Africa Middle Jurassic Africa Jurassic Madagascar Bajocian Stage Bathonian Stage Mudstone formations Limestone formations Lagoonal deposits Reef deposits Ichnofossiliferous formations Paleontology in Madagascar Formations