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Beech (''Fagus'') is a genus of deciduous trees in the family
Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as evergre ...
, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engleriana'' subgenus is found only in East Asia, distinctive for its low branches, often made up of several major trunks with yellowish bark. The better known ''Fagus'' subgenus beeches are high-branching with tall, stout trunks and smooth silver-grey bark. The European beech ('' Fagus sylvatica'') is the most commonly cultivated. Beeches are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
, bearing both male and female flowers on the same plant. The small flowers are unisexual, the female flowers borne in pairs, the male flowers wind-pollinating
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s. They are produced in spring shortly after the new leaves appear. The fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from the tree in autumn. They are small, roughly triangular, and edible, with a bitter, astringent, or mild and nut-like taste. The European species ''Fagus sylvatica'' yields a utility timber that is tough but dimensionally unstable. It is widely used for furniture framing and carcase construction, flooring and engineering purposes, in plywood, and household items like plates, but rarely as a decorative wood. The timber can be used to build chalets, houses, and log cabins. Beechwood also makes excellent
firewood Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form, compared to other forms of wood fuel like pellets or chips. Firewood can ...
, easily split and burning for many hours with bright but calm flames. Slats of washed beech wood are spread around the bottom of fermentation tanks for Budweiser beer. Beech logs are burned to dry the
malt Malt is germinated cereal grain that has been dried in a process known as " malting". The grain is made to germinate by soaking in water and is then halted from germinating further by drying with hot air. Malted grain is used to make beer, wh ...
used in some German smoked beers. Beech is also used to smoke Westphalian ham,
andouille Andouille ( , ; ; from Latin'induco') is a smoked sausage made using pork, originating in France. France In France, particularly Brittany and Normandy, the traditional ingredients of andouille are primarily pig chitterlings, tripe, onions, w ...
sausage, and some cheeses.


Description

The European beech ('' Fagus sylvatica'') is the most commonly cultivated, although few important differences are seen between species aside from detail elements such as leaf shape. The leaves of beech trees are entire or sparsely toothed, from long and broad. Beeches are
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
, bearing both male and female flowers on the same plant. The small flowers are unisexual, the female flowers borne in pairs, the male flowers wind-pollinating
catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...
s. They are produced in spring shortly after the new leaves appear. The bark is smooth and light gray. The fruit is a small, sharply three-angled nut long, borne singly or in pairs in soft-spined husks long, known as cupules. The husk can have a variety of spine- to scale-like appendages, the character of which is, in addition to leaf shape, one of the primary ways beeches are differentiated. The nuts have a bitter taste (though not nearly as bitter as
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...
s) and a high tannin content; these are called beechnuts or beech mast.


Taxonomy

Recent classification systems of the genus recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engleriana'' subgenus is found only in East Asia, and is notably distinct from the ''Fagus'' subgenus in that these beeches are low-branching trees, often made up of several major trunks with yellowish bark. Further differentiating characteristics include the whitish bloom on the underside of the leaves, the visible tertiary leaf veins, and a long, smooth cupule-peduncle. Proposed by botanist Chung-Fu Shen in 1992, '' F. japonica'', '' F. engleriana'', and '' F. okamotoi'' comprise this subgenus. The better known ''Fagus'' subgenus beeches are high-branching with tall, stout trunks and smooth silver-gray bark. This group includes ''F. sylvatica'', '' F. grandifolia'', '' F. crenata'', '' F. lucida'', '' F. longipetiolata'', and '' F. hayatae''. The classification of the European beech, ''F. sylvatica'', is complex, with a variety of different names proposed for different species and subspecies within this region (for example '' F. taurica'', '' F. orientalis'', and ''F. moesica''). Research suggests that beeches in Eurasia differentiated fairly late in evolutionary history, during the Miocene. The populations in this area represent a range of often overlapping morphotypes, and genetic analysis does not clearly support separate species. ''Fagus'' is the most basal group in the evolution of the
Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as evergre ...
family, which also includes oaks and
chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrelat ...
s. The southern beeches (genus '' Nothofagus'') previously thought closely related to beeches, are now treated as members of a separate family, the Nothofagaceae (which remains a member of the order Fagales). They are found in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, New Zealand, New Guinea,
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, and Argentina and Chile (principally Patagonia and Tierra del Fuego).


Species

The following is a partial list of taxa, which have been accepted as species at one point:


Natural Hybrids


Fossils

*†''
Fagus langevinii ''Fagus langevinii'' is an extinct species of beech in the family Fagaceae. The species is known from fossil fruits, nuts, pollen, and leaves found in the early Eocene deposits of South central British Columbia, and northern Washington state, ...
'' *†''
Fagus subferruginea Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...
''


Etymology

The name of the tree in Latin, ''fagus'' (from whence the generic epithet), is cognate with English "beech" and of Indo-European origin, and played an important role in early debates on the geographical origins of the Indo-European people, the
beech argument The beech argument (german: Das Buchenargument) is a now mostly outdated argument in Indo-European studies that is in favour of placing the Indo-European urheimat in an area west of a line connecting Kaliningrad and the Black Sea, based on the curr ...
. Greek φηγός (figós) is from the same root, but the word was transferred to the oak tree (e.g. Iliad 16.767) as a result of the absence of beech trees in southern Greece.


Distribution and habitat


Britain and Ireland

'' Fagus sylvatica'' was a late entrant to Great Britain after the last glaciation, and may have been restricted to basic soils in the south of England. Some suggest that it was introduced by Neolithic tribes who planted the trees for their edible nuts. The beech is classified as a native in the south of England and as a non-native in the north where it is often removed from 'native' woods. Large areas of the Chilterns are covered with beech woods, which are habitat to the common bluebell and other flora. The Cwm Clydach National Nature Reserve in southeast Wales was designated for its beech woodlands, which are believed to be on the western edge of their natural range in this steep limestone gorge. Beech is not native to Ireland; however, it was widely planted in the 18th century and can become a problem shading out the native woodland understory. Today, beech is widely planted for hedging and in deciduous woodlands, and mature, regenerating stands occur throughout mainland Britain at elevations below about . The tallest and longest hedge in the world (according to '' Guinness World Records'') is the Meikleour Beech Hedge in Meikleour,
Perth and Kinross Perth and Kinross ( sco, Pairth an Kinross; gd, Peairt agus Ceann Rois) is one of the 32 council areas of Scotland and a Lieutenancy Area. It borders onto the Aberdeenshire, Angus, Argyll and Bute, Clackmannanshire, Dundee, Fife, Highland and S ...
, Scotland.


Continental Europe

''Fagus sylvatica'' is one of the most common hardwood trees in north-central Europe, in France constituting alone about 15% of all nonconifers. Eastern Europe is also home to the lesser-known oriental beech (''F. orientalis'') and Crimean beech (''F. taurica''). As a naturally growing forest tree, beech marks the important border between the European deciduous forest zone and the northern pine forest zone. This border is important for wildlife and fauna. In Denmark and Scania at the southernmost peak of the Scandinavian peninsula, southwest of the natural
spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
boundary, it is the most common forest tree. It grows naturally in Denmark and southern Norway and Sweden up to about 57–59°N. The most northern known naturally growing (not planted) beech trees are found in a small grove north of Bergen on the west coast of Norway. Near the city of
Larvik Larvik () is a List of cities in Norway, town and Municipalities of Norway, municipality in Vestfold in Vestfold og Telemark Counties of Norway, county, Norway. The administrative centre of the municipality is the city of Larvik. The municipality ...
is the largest naturally occurring beech forest in Norway,
Bøkeskogen Bøkeskogen ("The Beech Tree Forest") stretches inwards along Farris, Farris Lake in Larvik. It is an archeological area which is home to between 83, 90, and 100 burial mounds.Bertelsen, Hans Kristian (1998). ''Bli kjent med Vestfold / Become acqua ...
. Some research suggests that early agriculture patterns supported the spread of beech in continental Europe. Research has linked the establishment of beech stands in Scandinavia and Germany with cultivation and fire disturbance, i.e. early agricultural practices. Other areas which have a long history of cultivation, Bulgaria for example, do not exhibit this pattern, so how much human activity has influenced the spread of beech trees is as yet unclear. The
primeval beech forests of the Carpathians Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe is a transnational serial nature UNESCO World Heritage Site, encompassing 94 component parts (forests of European beech, ''Fagus sylvatica'') in 18 European countr ...
are also an example of a singular, complete, and comprehensive forest dominated by a single tree species - the beech tree. Forest dynamics here were allowed to proceed without interruption or interference since the last ice age. Nowadays, they are amongst the last pure beech forests in Europe to document the undisturbed postglacial repopulation of the species, which also includes the unbroken existence of typical animals and plants. These virgin beech forests and similar forests across 12 countries in continental Europe were inscribed on the UNESCO World Heritage List in 2007.


North America

The American beech (''Fagus grandifolia'') occurs across much of the eastern United States and southeastern Canada, with a disjunct population in Mexico. It is the only ''Fagus'' species in the Western Hemisphere. Before the Pleistocene Ice Age, it is believed to have spanned the entire width of the continent from the Atlantic Ocean to the Pacific but now is confined to the east of the Great Plains. ''F. grandifolia'' tolerates hotter climates than European species but is not planted much as an ornamental due to slower growth and less resistance to urban pollution. It most commonly occurs as an overstory component in the northern part of its range with sugar maple, transitioning to other forest types further south such as beech-magnolia. American beech is rarely encountered in developed areas except as a remnant of a forest that was cut down for land development. The dead brown leaves of the American beech remain on the branches until well into the following spring, when the new buds finally push them off.


Asia

East Asia is home to five species of ''Fagus'', only one of which (''F. crenata'') is occasionally planted in Western countries. Smaller than ''F. sylvatica'' and ''F. grandifolia'', this beech is one of the most common hardwoods in its native range.


Ecology

Beech grows on a wide range of soil types, acidic or basic, provided they are not waterlogged. The tree canopy casts dense shade and thickens the ground with leaf litter. In North America, they can form beech-maple
climax Climax may refer to: Language arts * Climax (narrative), the point of highest tension in a narrative work * Climax (rhetoric), a figure of speech that lists items in order of importance Biology * Climax community, a biological community th ...
forests by partnering with the sugar maple. The
beech blight aphid The beech blight aphid (''Grylloprociphilus imbricator'') is a small insect in the order Hemiptera that feed primarily on the sap of Fagus grandifolia, American beech trees. The aphids form dense colonies on small branches and the undersides of l ...
(''Grylloprociphilus imbricator'') is a common pest of American beech trees. Beeches are also used as
food plants A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponics ...
by some species of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
. Beech bark is extremely thin and scars easily. Since the beech tree has such delicate bark, carvings, such as lovers' initials and other forms of graffiti, remain because the tree is unable to heal itself.


Diseases

Beech bark disease is a fungal infection that attacks the American beech through damage caused by scale insects. Infection can lead to the death of the tree.
Beech leaf disease ''Beech leaf disease'' is a newly discovered lethal disease of beech trees believed to be caused by the nematode '' Litylenchus crenatae mccannii''. The symptoms of the disease appear as a dark green, interveinal banding pattern on the lower canopy ...
is a disease spread by the newly discovered nematode, ''Litylenchus crenatae mccannii''. This disease was first discovered in Lake County, Ohio, in 2012 and has now spread to over 41 counties in Ohio, Pennsylvania, New York, and Ontario, Canada.


Cultivation

The beech most commonly grown as an
ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
is the European beech (''Fagus sylvatica''), widely cultivated in North America and its native Europe. Many varieties are in cultivation, notably the weeping beech ''F. sylvatica'' 'Pendula', several varieties of copper or purple beech, the fern-leaved beech ''F. sylvatica'' 'Asplenifolia', and the tricolour beech ''F. sylvatica'' 'Roseomarginata'. The columnar Dawyck beech (''F. sylvatica'' 'Dawyck') occurs in green, gold, and purple forms, named after Dawyck Botanic Garden in the Scottish Borders, one of the four garden sites of the Royal Botanic Garden Edinburgh.


Uses

Beech wood is an excellent
firewood Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form, compared to other forms of wood fuel like pellets or chips. Firewood can ...
, easily split and burning for many hours with bright but calm flames. Slats of beech wood are washed in caustic soda to leach out any flavour or aroma characteristics and are spread around the bottom of fermentation tanks for Budweiser beer. This provides a complex surface on which the yeast can settle, so that it does not pile up, preventing yeast autolysis which would contribute off-flavours to the beer. Beech logs are burned to dry the
malt Malt is germinated cereal grain that has been dried in a process known as " malting". The grain is made to germinate by soaking in water and is then halted from germinating further by drying with hot air. Malted grain is used to make beer, wh ...
used in German smoked beers. Beech is also used to smoke Westphalian ham, traditional
andouille Andouille ( , ; ; from Latin'induco') is a smoked sausage made using pork, originating in France. France In France, particularly Brittany and Normandy, the traditional ingredients of andouille are primarily pig chitterlings, tripe, onions, w ...
(an offal sausage) from Normandy, and some cheeses. Some
drum The drum is a member of the percussion group of musical instruments. In the Hornbostel-Sachs classification system, it is a membranophone. Drums consist of at least one membrane, called a drumhead or drum skin, that is stretched over a she ...
s are made from beech, which has a tone between those of maple and birch, the two most popular drum woods. The textile modal is a kind of
rayon Rayon is a semi-synthetic fiber, made from natural sources of regenerated cellulose, such as wood and related agricultural products. It has the same molecular structure as cellulose. It is also called viscose. Many types and grades of viscose f ...
often made wholly from reconstituted cellulose of pulped beech wood. The European species ''Fagus sylvatica'' yields a utility timber that is tough but dimensionally unstable. It weighs about 720 kg per cubic metre and is widely used for furniture framing and carcase construction, flooring, and engineering purposes, in plywood and household items like plates, but rarely as a decorative wood. The timber can be used to build chalets, houses, and log cabins. Beech wood is used for the stocks of military rifles when traditionally preferred woods such as walnut are scarce or unavailable or as a lower-cost alternative. The edible fruit of the beech tree, known as beechnuts or mast, is found in small burrs that drop from the tree in autumn. They are small, roughly triangular, and edible, with a bitter, astringent, or in some cases, mild and nut-like taste. According to the Roman statesman Pliny the Elder in his work ''Natural History'', beechnut was eaten by the people of Chios when the town was besieged, writing of the fruit: "that of the beech is the sweetest of all; so much so, that, according to Cornelius Alexander, the people of the city of Chios, when besieged, supported themselves wholly on mast". The leaves can be steeped in liquor to give a light green/yellow liqueur. They can also be roasted and pulverized into an adequate coffee substitute. In antiquity, the bark of the beech tree was used by Indo-European people for writing-related purposes, especially in a religious context. Beech wood tablets were a common writing material in Germanic societies before the development of paper. The Old English ''bōc'' has the primary sense of "beech" but also a secondary sense of "book", and it is from ''bōc'' that the modern word derives. In modern German, the word for "book" is ''Buch,'' with ''Buche'' meaning "beech tree". In modern Dutch, the word for "book" is ''boek,'' with ''beuk'' meaning "beech tree". In Swedish, these words are the same, ''bok'' meaning both "beech tree" and "book". There is a similar relationship in some Slavic languages. In Russian and Bulgarian, the word for beech is бук (''buk''), while that for "letter" (as in a letter of the alphabet) is буква (''bukva''), while Serbo-Croatian and Slovene use " bukva" to refer to the tree. The pigment bistre was made from beech wood soot. Beech litter raking as a replacement for straw in
animal husbandry Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, fibre, milk, or other products. It includes day-to-day care, selective breeding, and the raising of livestock. Husbandry has a long history, starti ...
was an old non-timber practice in forest management that once occurred in parts of
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
in the 17th century. Beech has been listed as one of the 38 plants whose flowers are used to prepare
Bach flower remedies Bach flower remedies (BFRs) are solutions of brandy and water—the water containing extreme dilutions of flower material developed by Edward Bach, an English homeopath, in the 1930s. Bach claimed that the dew found on flower petals retains t ...
.


See also

*
Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe Ancient and Primeval Beech Forests of the Carpathians and Other Regions of Europe is a transnational serial nature UNESCO World Heritage Site, encompassing 94 component parts (forests of European beech, ''Fagus sylvatica'') in 18 European countr ...
* English Lowlands beech forests * The Weeping Beech


References


External links

* *
Traditional and Modern Use of Beech
{{Authority control Edible nuts and seeds Ornamental trees