Bears are
carnivora
Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
n
mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s of the
family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Ursidae. They are classified as
caniforms
Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group. The ...
, or doglike carnivorans. Although only eight
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of bears are extant, they are widespread, appearing in a wide variety of
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found on the continents of North America, South America, Europe, and Asia. Common characteristics of modern bears include large bodies with stocky legs, long snouts, small rounded ears, shaggy hair,
plantigrade
151px, Portion of a human skeleton, showing plantigrade habit
In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...
paws with five nonretractile claws, and short tails.
While the
polar bear
The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
is mostly
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
, and the
giant panda
The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear (or simply the panda), is a bear species endemic to China. It is characterised by its bold black-and-white coat and rotund body. The name "giant panda" is sometimes us ...
feeds almost entirely on
bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
, the remaining six species are
omnivorous
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...
with varied diets. With the exception of
courting
Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
individuals and mothers with their young, bears are typically
solitary animals
Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies.
Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
. They may be
diurnal or
nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.
Nocturnal creatures generally have highly developed sens ...
and have an excellent
sense of smell
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste.
In humans, it ...
. Despite their heavy build and awkward gait, they are adept runners, climbers, and swimmers. Bears use shelters, such as caves and logs, as their dens; most species occupy their dens during the winter for a long period of
hibernation
Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...
, up to 100 days.
Bears have been hunted since
prehistoric
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
times for
their meat and fur; they have been used for
bear-baiting
Bear-baiting is a blood sport in which a chained bear and one or more dogs are forced to fight one another. It may also involve pitting a bear against another animal.
History Europe Great Britain
Bear-baiting was very popular from the 12th ...
and other forms of entertainment, such as being
made to dance. With their powerful physical presence, they play a prominent role in
the arts
The arts are a very wide range of human practices of creative expression, storytelling and cultural participation. They encompass multiple diverse and plural modes of thinking, doing and being, in an extremely broad range of media. Both ...
,
mythology
Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narrat ...
, and other cultural aspects of various human societies. In modern times, bears have come under pressure through encroachment on their habitats and illegal trade in bear parts, including the Asian
bile bear
Bile bears, sometimes called battery bears, are bears kept in captivity to harvest their bile, a digestive fluid produced by the liver and stored in the gallbladder, which is used by some traditional Asian medicine practitioners. It is estimated t ...
market. The
IUCN
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
lists six bear species as
vulnerable or
endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
, and even
least concern
A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
species, such as the
brown bear
The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
, are at risk of
extirpation
Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
in certain countries. The poaching and international trade of these most threatened populations are prohibited, but still ongoing.
Etymology
The English word "bear" comes from
Old English
Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
' and belongs to a family of names for the bear in
Germanic languages
The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania and Southern Africa. The most widely spoken Germanic language, Engli ...
, such as Swedish ', also used as a first name. This form is conventionally said to be related to a
Proto-Indo-European
Proto-Indo-European (PIE) is the reconstructed common ancestor of the Indo-European language family. Its proposed features have been derived by linguistic reconstruction from documented Indo-European languages. No direct record of Proto-Indo-E ...
word for "brown", so that "bear" would mean "the brown one". However,
Ringe notes that while this etymology is semantically plausible, a word meaning "brown" of this form cannot be found in Proto-Indo-European. He suggests instead that "bear" is from the Proto-Indo-European word ''*ǵʰwḗr- ~ *ǵʰwér'' "wild animal". This terminology for the animal originated as a
taboo avoidance term: proto-Germanic tribes replaced their original word for bear—''arkto''—with this euphemistic expression out of fear that speaking the animal's true name might cause it to appear. According to author
Ralph Keyes
Ralph (pronounced ; or ,) is a male given name of English, Scottish and Irish origin, derived from the Old English ''Rædwulf'' and Radulf, cognate with the Old Norse ''Raðulfr'' (''rað'' "counsel" and ''ulfr'' "wolf").
The most common forms ...
, this is the oldest known
euphemism
A euphemism () is an innocuous word or expression used in place of one that is deemed offensive or suggests something unpleasant. Some euphemisms are intended to amuse, while others use bland, inoffensive terms for concepts that the user wishes ...
.
Bear taxon names such as
Arctoidea
Arctoidea is a clade of mostly carnivorous mammals which include the extinct Hemicyonidae (dog-bears), and the extant Musteloidea (weasels, raccoons, skunks, red pandas), Pinnipedia (seals, sea lions), and Ursidae (bears), found in all continents ...
and ''
Helarctos
The sun bear (''Helarctos malayanus'') is a species in the family Ursidae (the only species in the genus ''Helarctos'') occurring in the tropical forests of Southeast Asia. It is the smallest bear, standing nearly at the shoulder and weighing ...
'' come from the ancient Greek ἄρκτος (''arktos''), meaning bear, as do the names "
arctic
The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
" and "
antarctic
The Antarctic ( or , American English also or ; commonly ) is a polar region around Earth's South Pole, opposite the Arctic region around the North Pole. The Antarctic comprises the continent of Antarctica, the Kerguelen Plateau and other ...
", via the name of the
constellation
A constellation is an area on the celestial sphere in which a group of visible stars forms Asterism (astronomy), a perceived pattern or outline, typically representing an animal, mythological subject, or inanimate object.
The origins of the e ...
Ursa Major
Ursa Major (; also known as the Great Bear) is a constellation in the northern sky, whose associated mythology likely dates back into prehistory. Its Latin name means "greater (or larger) bear," referring to and contrasting it with nearby Ursa ...
, the "Great Bear", prominent in the northern sky.
Bear taxon names such as Ursidae and ''Ursus'' come from Latin ''Ursus/Ursa'', he-bear/she-bear.
[ The female first name " Ursula", originally derived from a Christian ]saint
In religious belief, a saint is a person who is recognized as having an exceptional degree of Q-D-Š, holiness, likeness, or closeness to God. However, the use of the term ''saint'' depends on the context and Christian denomination, denominat ...
's name, means "little she-bear" (diminutive of Latin ''ursa''). In Switzerland, the male first name "Urs" is especially popular, while the name of the canton and city of Bern
german: Berner(in)french: Bernois(e) it, bernese
, neighboring_municipalities = Bremgarten bei Bern, Frauenkappelen, Ittigen, Kirchlindach, Köniz, Mühleberg, Muri bei Bern, Neuenegg, Ostermundigen, Wohlen bei Bern, Zollikofen
, website ...
is derived from ''Bär'', German for bear. The Germanic name Bernard
Bernard (''Bernhard'') is a French and West Germanic masculine given name. It is also a surname.
The name is attested from at least the 9th century. West Germanic ''Bernhard'' is composed from the two elements ''bern'' "bear" and ''hard'' "brav ...
(including Bernhardt and similar forms) means "bear-brave", "bear-hardy", or "bold bear". The Old English name Beowulf
''Beowulf'' (; ang, Bēowulf ) is an Old English epic poem in the tradition of Germanic heroic legend consisting of 3,182 alliterative lines. It is one of the most important and most often translated works of Old English literature. The ...
is a kenning
A kenning ( Icelandic: ) is a figure of speech in the type of circumlocution, a compound that employs figurative language in place of a more concrete single-word noun. Kennings are strongly associated with Old Norse-Icelandic and Old English po ...
, "bee-wolf", for bear, in turn meaning a brave warrior.
Taxonomy
The family Ursidae is one of nine families in the suborder Caniformia
Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include dogs (wolves, foxes, etc.), bears, raccoons, and mustelids. The Pinnipedia (seals, walruses and sea lions) are also assigned to this group. ...
, or "doglike" carnivorans, within the order
Order, ORDER or Orders may refer to:
* Categorization, the process in which ideas and objects are recognized, differentiated, and understood
* Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Carnivora
Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
. Bears' closest living relatives are the pinniped
Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walru ...
s, canids
Canidae (; from Latin, ''canis'', "dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within th ...
, and musteloids. Modern bears comprise eight species in three subfamilies: Ailuropodinae
Ailuropodinae is a subfamily of Ursidae that contains only one extant species, the giant panda (''Ailuropoda melanoleuca'') of China. The fossil record of this group has shown that various species of pandas were more widespread across the Holarct ...
(monotypic with the giant panda
The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear (or simply the panda), is a bear species endemic to China. It is characterised by its bold black-and-white coat and rotund body. The name "giant panda" is sometimes us ...
), Tremarctinae
The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear (''Tremarctos ornatus'') of South America, and several extinct species from four genera: the Tremarctos floridanus, Florid ...
(monotypic with the spectacled bear), and Ursinae
Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998) and Jin et al. (2007).
Classification
The genera ''Melursus'' and ''Helarctos'' are sometimes also included in ''Ursus'' ...
(containing six species divided into one to three genera, depending on the authority). Nuclear chromosome analysis show that the karyotype
A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
of the six ursine bears is nearly identical, each having 74 chromosome
A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s (''see'' Ursid hybrid
An ursid hybrid is an animal with parents from two different species or subspecies of the bear family (''Ursidae''). Species and subspecies of bear known to have produced offspring with another bear species or subspecies include black bears, gri ...
), whereas the giant panda has 42 chromosomes and the spectacled bear 52. These smaller numbers can be explained by the fusing of some chromosomes, and the banding patterns on these match those of the ursine species, but differ from those of procyonids, which supports the inclusion of these two species in Ursidae rather than in Procyonidae
Procyonidae is a New World family of the order Carnivora. It comprises the raccoons, ringtails, cacomistles, coatis, kinkajous, olingos, and olinguitos. Procyonids inhabit a wide range of environments and are generally omnivorous.
Character ...
, where they had been placed by some earlier authorities.
Evolution
The earliest members of Ursidae belong to the extinct subfamily Amphicynodontinae, including ''Parictis
''Parictis'' is an extinct arctoid, possibly the earliest genus of bears known, though a recent variety of morphological evidence links amphicynodontines with pinnipeds. It was a very small and graceful arctoid with a skull only 7 cm long. ...
'' (late Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
to early middle Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, 38–18 Mya) and the slightly younger ''Allocyon'' (early Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
, 34–30 Mya), both from North America. These animals looked very different from today's bears, being small and raccoon
The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
-like in overall appearance, with diets perhaps more similar to that of a badger
Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united b ...
. ''Parictis'' does not appear in Eurasia and Africa until the Miocene. It is unclear whether late-Eocene ursids were also present in Eurasia, although faunal exchange across the Bering land bridge
Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of ...
may have been possible during a major sea level low stand as early as the late Eocene (about 37 Mya) and continuing into the early Oligocene. European genera morphologically very similar to ''Allocyon'', and to the much younger American ''Kolponomos
''Kolponomos'' is an extinct genus of carnivoran mammal that existed in the Late Arikareean North American Land Mammal Age, early Miocene epoch, about 20 million years ago. It was likely a marine mammal. The genus was erected in 1960 by Ruben A. ...
'' (about 18 Mya), are known from the Oligocene, including ''Amphicticeps'' and ''Amphicynodon
''Amphicynodon'' was an extinct genus of caniform carnivore. It has traditionally been considered early bear, although recent evidence has suggested it may be a unique member linked to other pinnipeds. It was endemic to Europe and Asia during t ...
''.[ There has been various morphological evidence linking amphicynodontines with ]pinniped
Pinnipeds (pronounced ), commonly known as seals, are a widely distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant families Odobenidae (whose only living member is the walru ...
s, as both groups were semi-aquatic, otter-like mammals. In addition to the support of the pinniped–amphicynodontine clade, other morphological and some molecular evidence supports bears being the closest living relatives to pinnipeds.[
]
The raccoon-sized, dog-like ''Cephalogale
''Cephalogale'' is an extinct genus of hemicyonine bear which lived in the Oligocene and Early Miocene epochs in North America and Europe. It lived from around 28.4—20.0 Mya. Before it was reconsidered to be close to the ancestry of hemicyo ...
'' is the oldest-known member of the subfamily Hemicyoninae
Hemicyoninae is an extinct subfamily of Ursidae, often called dog bears (literally "half dog" (Ancient Greek, Greek: )). They were bear-like carnivorans living in Europe, North America, Africa and Asia during the Oligocene through Miocene epoch ...
, which first appeared during the middle Oligocene in Eurasia about 30 Mya.[ The subfamily includes the younger genera '']Phoberocyon
''Phoberocyon'' is a large extinct genus of hemicyonine bear, found primarily in North America during the Miocene. It lived from 28.4 to 13.7 mya, existing for approximately . One species, ''P. hispanicus'', is known from Miocene Spain
, im ...
'' (20–15 Mya), and ''Plithocyon
''Plithocyon'' is an extinct genus of hemicyonine bear of the Miocene epoch, endemic to North America and Europe. It lived from ~15.97—11.61 Ma, existing for approximately .
Fossil distribution
Sites and age of some specimens:
*Hemicyon Qua ...
'' (15–7 Mya). A ''Cephalogale''-like species gave rise to the genus ''Ursavus
''Ursavus'' is an extinct genus of bear that existed in North America, Europe, and Asia during the Miocene period, about 23–5.3 million years ago (Mya), existing for roughly . The genus apparently dispersed from Asia into North America abo ...
'' during the early Oligocene (30–28 Mya); this genus proliferated into many species in Asia and is ancestral to all living bears. Species of ''Ursavus'' subsequently entered North America, together with ''Amphicynodon'' and ''Cephalogale'', during the early Miocene (21–18 Mya). Members of the living lineages of bears diverged from ''Ursavus'' between 15 and 20 Mya, likely via the species ''Ursavus elmensis''. Based on genetic and morphological data, the Ailuropodinae (pandas) were the first to diverge from other living bears about 19 Mya, although no fossils of this group have been found before about 11 Mya.
The New World short-faced bears (Tremarctinae) differentiated from Ursinae following a dispersal event into North America during the mid-Miocene (about 13 Mya).[ They invaded South America (≈2.5 or 1.2 Ma) following formation of the ]Isthmus of Panama
The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
. Their earliest fossil representative is ''Plionarctos
''Plionarctos'' is an extinct genus of bear endemic to North America from the Miocene to the Pliocene, ~10.3—3.3 Mya (unit), Mya, existing for about 7 million years.
''Indarctos'' (10.7—9.2 Mya) preceded ''Plionarctos'' by only a few thousan ...
'' in North America (c. 10–2 Ma). This genus is probably the direct ancestor to the North American short-faced bears (genus ''Arctodus''), the South American short-faced bears (''Arctotherium''), and the spectacled bears, ''Tremarctos'', represented by both an extinct North American species ('' T. floridanus''), and the lone surviving representative of the Tremarctinae, the South American spectacled bear (''T. ornatus'').[
]
The subfamily Ursinae experienced a dramatic proliferation of taxa about 5.3–4.5 Mya, coincident with major environmental changes; the first members of the genus ''Ursus
Ursus is Latin for bear. It may also refer to:
Animals
* ''Ursus'' (mammal), a genus of bears
People
* Ursus of Aosta, 6th-century evangelist
* Ursus of Auxerre, 6th-century bishop
* Ursus of Solothurn, 3rd-century martyr
* Ursus (''praefectus ...
'' appeared around this time. The sloth bear
The sloth bear (''Melursus ursinus'') is a myrmecophagous bear species native to the Indian subcontinent. It feeds on fruits, ants and termites. It is listed as Vulnerable species, vulnerable on the IUCN Red List, mainly because of habitat loss ...
is a modern survivor of one of the earliest lineages to diverge during this radiation event (5.3 Mya); it took on its peculiar morphology, related to its diet of termites and ants, no later than by the early Pleistocene. By 3–4 Mya, the species ''Ursus minimus'' appears in the fossil record of Europe; apart from its size, it was nearly identical to today's Asian black bear
The Asian black bear (''Ursus thibetanus''), also known as the Asiatic black bear, moon bear and white-chested bear, is a medium-sized bear species native to Asia that is largely adapted to an arboreal lifestyle. It lives in the Himalayas, sout ...
. It is likely ancestral to all bears within Ursinae, perhaps aside from the sloth bear. Two lineages evolved from ''U. minimus'': the black bears (including the sun bear
The sun bear (''Helarctos malayanus'') is a species in the family Ursidae (the only species in the genus ''Helarctos'') occurring in the tropical forests of Southeast Asia. It is the smallest bear, standing nearly at the shoulder and weighing ...
, the Asian black bear, and the American black bear
The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
); and the brown bears (which includes the polar bear
The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
). Modern brown bear
The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
s evolved from ''U. minimus'' via ''Ursus etruscus
''Ursus etruscus'' (the Etruscan bear) is an extinct species of bear, endemic to Europe, Asia and North Africa during the Pliocene through Pleistocene, living from ~5.3 million to 100,000 years ago.
Taxonomy
''Ursus etruscus'' appears ...
'', which itself is ancestral to the extinct Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
cave bear
The cave bear (''Ursus spelaeus'') is a prehistoric species of bear that lived in Europe and Asia during the Pleistocene and became extinct about 24,000 years ago during the Last Glacial Maximum.
Both the word "cave" and the scientific name ' ...
.[ Species of Ursinae have migrated repeatedly into North America from Eurasia as early as 4 Mya during the early Pliocene.] The polar bear is the most recently evolved species and descended from a population of brown bears that became isolated in northern latitudes by glaciation 400,000 years ago.
Phylogeny
The relationship of the bear family with other carnivorans is shown in the following phylogenetic tree
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
, which is based on the molecular phylogenetic analysis of six genes in Flynn, 2005.
Note that although they are called "bears" in some languages, red panda
The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
s and raccoons and their close relatives are not bears, but rather musteloids.[
There are two phylogenetic hypotheses on the relationships among extant and fossil bear species. One is all species of bears are classified in seven subfamilies as adopted here and related articles: ]Amphicynodontinae
Amphicynodontidae is a probable clade of extinct arctoids. While some researchers consider this group to be an extinct subfamily of bears, a variety of morphological evidence links amphicynodontines with pinnipeds, as the group were semi-aquati ...
, Hemicyoninae
Hemicyoninae is an extinct subfamily of Ursidae, often called dog bears (literally "half dog" (Ancient Greek, Greek: )). They were bear-like carnivorans living in Europe, North America, Africa and Asia during the Oligocene through Miocene epoch ...
, Ursavinae
Ursavini is an extinct tribe of mammals of the family Ursidae (bears) endemic to North America
North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by ...
, Agriotheriinae
''Agriotherium'' is an extinct genus of bears whose fossils are found in Miocene through Pleistocene-aged strata of North America, Europe, Africa, and Asia. This long-lived genus persisted from at least ~11.6–2.5 Mya. Materials from the late-s ...
, Ailuropodinae
Ailuropodinae is a subfamily of Ursidae that contains only one extant species, the giant panda (''Ailuropoda melanoleuca'') of China. The fossil record of this group has shown that various species of pandas were more widespread across the Holarct ...
, Tremarctinae
The Tremarctinae or short-faced bears is a subfamily of Ursidae that contains one living representative, the spectacled bear (''Tremarctos ornatus'') of South America, and several extinct species from four genera: the Tremarctos floridanus, Florid ...
, and Ursinae
Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998) and Jin et al. (2007).
Classification
The genera ''Melursus'' and ''Helarctos'' are sometimes also included in ''Ursus'' ...
. Below is a cladogram of the subfamilies of bears after McLellan and Reiner (1992)[ and Qiu ''et al''. (2014):]
The second alternative phylogenetic hypothesis was implemented by McKenna et al. (1997) to classify all the bear species into the superfamily Ursoidea, with Hemicyoninae and Agriotheriinae being classified in the family "Hemicyonidae". Amphicynodontinae under this classification were classified as stem-pinnipeds
Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammal, marine mammals. They comprise the extant taxon, extant family (biology ...
in the superfamily Phocoidea. In the McKenna and Bell classification both bears and pinnipeds are in a parvorder of carnivora
Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
n mammals known as Ursida, along with the extinct bear dogs of the family Amphicyonidae
Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene (around 45 mya), spread to Europe by the late Eocene (35 mya), and appear in Asia, ...
. Below is the cladogram based on McKenna and Bell (1997) classification:
The phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
of extant bear species is shown in a cladogram based on complete mitochondrial DNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
sequences from Yu ''et al''. (2007) The giant panda, followed by the spectacled bear, are clearly the oldest species. The relationships of the other species are not very well resolved, though the polar bear and the brown bear form a close grouping.[
]
Physical characteristics
Size
The bear family includes the most massive extant terrestrial members of the order Carnivora. The polar bear is considered to be the largest extant species, with adult males weighing and measuring in total length. The smallest species is the sun bear, which ranges in weight and in length. Prehistoric North
North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography.
Etymology
T ...
and South American short-faced bear
''Arctotherium'' ("''bear beast''") is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. ''Arctotherium'' migrated from North America to South America during the Great American Interchange, following the f ...
s were the largest species known to have lived. The latter estimated to have weighed and stood tall. Body weight varies throughout the year in bears of temperate and arctic climates, as they build up fat reserves in the summer and autumn and lose weight during the winter.
Morphology
Bears are generally bulky and robust animals with short tails. They are sexually dimorphic
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
with regard to size, with males typically being larger. Larger species tend to show increased levels of sexual dimorphism in comparison to smaller species.[ Relying as they do on strength rather than speed, bears have relatively short limbs with thick bones to support their bulk. The shoulder blades and the ]pelvis
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton).
The ...
are correspondingly massive. The limbs are much straighter than those of the big cat
The term "big cat" is typically used to refer to any of the five living members of the genus '' Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard.
Despite enormous differences in size, various cat species are quite similar ...
s as there is no need for them to flex in the same way due to the differences in their gait. The strong forelimbs are used to catch prey, to excavate dens, to dig out burrowing animals, to turn over rocks and logs to locate prey, and to club large creatures.
Unlike most other land carnivorans, bears are plantigrade
151px, Portion of a human skeleton, showing plantigrade habit
In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...
. They distribute their weight toward the hind feet, which makes them look lumbering when they walk. They are capable of bursts of speed but soon tire, and as a result mostly rely on ambush rather than the chase. Bears can stand on their hind feet and sit up straight with remarkable balance. Their front paws are flexible enough to grasp fruit and leaves. Bears' non-retractable claws are used for digging, climbing, tearing, and catching prey. The claws on the front feet are larger than those on the back and may be a hindrance when climbing trees; black bears are the most arboreal of the bears, and have the shortest claws. Pandas are unique in having a bony extension on the wrist of the front feet which acts as a thumb, and is used for gripping bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
shoots as the animals feed.
Most mammals have agouti
The agouti (, ) or common agouti is any of several rodent species of the genus ''Dasyprocta''. They are native to Middle America, northern and central South America, and the southern Lesser Antilles. Some species have also been introduced else ...
hair, with each individual hair shaft having bands of color corresponding to two different types of melanin
Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
pigment. Bears however have a single type of melanin and the hairs have a single color throughout their length, apart from the tip which is sometimes a different shade. The coat consists of long guard hairs, which form a protective shaggy covering, and short dense hairs which form an insulating layer trapping air close to the skin. The shaggy coat helps maintain body heat during winter hibernation and is shed in the spring leaving a shorter summer coat. Polar bears have hollow, translucent guard hairs which gain heat from the sun and conduct it to the dark-colored skin below. They have a thick layer of blubber for extra insulation, and the soles of their feet have a dense pad of fur. While bears tend to be uniform in color, some species may have markings on the chest or face and the giant panda has a bold black-and-white pelage.
Bears have small rounded ears so as to minimize heat loss, but neither their hearing or sight are particularly acute. Unlike many other carnivorans they have color vision
Color vision, a feature of visual perception, is an ability to perceive differences between light composed of different wavelengths (i.e., different spectral power distributions) independently of light intensity. Color perception is a part of ...
, perhaps to help them distinguish ripe nuts and fruits. They are unique among carnivorans in not having touch-sensitive whiskers
Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to touch, sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are c ...
on the muzzle; however, they have an excellent sense of smell
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste.
In humans, it ...
, better than that of the dog, or possibly any other mammal. They use smell for signalling to each other (either to warn off rivals or detect mates) and for finding food. Smell is the principal sense used by bears to locate most of their food, and they have excellent memories which helps them to relocate places where they have found food before.
The skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
s of bears are massive, providing anchorage for the powerful masseter
In human anatomy, the masseter is one of the muscles of mastication. Found only in mammals, it is particularly powerful in herbivores to facilitate chewing of plant matter. The most obvious muscle of mastication is the masseter muscle, since it ...
and temporal jaw muscles. The canine teeth
In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened howeve ...
are large but mostly used for display, and the molar teeth flat and crushing. Unlike most other members of the Carnivora, bears have relatively undeveloped carnassial
Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
teeth, and their teeth are adapted for a diet that includes a significant amount of vegetable matter. Considerable variation occurs in dental formula
Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
even within a given species. This may indicate bears are still in the process of evolving from a mainly meat-eating diet to a predominantly herbivorous one. Polar bears appear to have secondarily re-evolved carnassial-like cheek teeth, as their diets have switched back towards carnivory. Sloth bears lack lower central incisors and use their protrusible lips for sucking up the termites on which they feed. The general dental formula for living bears is: . The structure of the larynx
The larynx (), commonly called the voice box, is an organ in the top of the neck involved in breathing, producing sound and protecting the trachea against food aspiration. The opening of larynx into pharynx known as the laryngeal inlet is about ...
of bears appears to be the most basal of the caniforms. They possess air pouches connected to the pharynx
The pharynx (plural: pharynges) is the part of the throat behind the mouth and nasal cavity, and above the oesophagus and trachea (the tubes going down to the stomach and the lungs). It is found in vertebrates and invertebrates, though its struc ...
which may amplify their vocalizations.
Bears have a fairly simple digestive system typical for carnivorans, with a single stomach, short undifferentiated intestines and no cecum
The cecum or caecum is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix (anatomy), appendix, to which i ...
. Even the herbivorous giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes. Its ability to digest cellulose
Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
is ascribed to the microbes in its gut. Bears must spend much of their time feeding in order to gain enough nutrition from foliage. The panda, in particular, spends 12–15 hours a day feeding.
Distribution and habitat
Extant bears are found in sixty countries primarily in the Northern Hemisphere and are concentrated in Asia, North America, and Europe. An exception is the spectacled bear; native to South America, it inhabits the Andean region. The sun bear
The sun bear (''Helarctos malayanus'') is a species in the family Ursidae (the only species in the genus ''Helarctos'') occurring in the tropical forests of Southeast Asia. It is the smallest bear, standing nearly at the shoulder and weighing ...
's range extends below the equator in Southeast Asia. The Atlas bear
The Atlas bear and African bearBryden, H. A. (ed.) (1899)''Great and small game of Africa''Rowland Ward Ltd., London. Pp. 544–608.#Schaller, Schaller, p. 220–21. (''Ursus arctos crowtheri'') is an extinct population or populations of brown be ...
, a subspecies of the brown bear was distributed in North Africa from Morocco to Libya, but it became extinct around the 1870s.
The most widespread species is the brown bear
The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
, which occurs from Western Europe eastwards through Asia to the western areas of North America. The American black bear
The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
is restricted to North America, and the polar bear
The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
is restricted to the Arctic Sea. All the remaining species of bear are Asian. They occur in a range of habitats which include tropical lowland rainforest, both coniferous and broadleaf forests, prairies, steppes, montane grassland, alpine scree slopes, Arctic tundra and in the case of the polar bear, ice floes
An ice floe () is a large pack ice, pack of Drift ice, floating ice often defined as a flat piece at least 20 m across at its widest point, and up to more than 10 km across. Drift ice is a floating field of sea ice composed of several ice fl ...
.[ Bears may dig their dens in hillsides or use caves, hollow logs and dense vegetation for shelter.][
]
Behavior and ecology
Brown and American black bears are generally diurnal, meaning that they are active for the most part during the day, though they may forage substantially by night. Other species may be nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.
Nocturnal creatures generally have highly developed sens ...
, active at night, though female sloth bears with cubs may feed more at daytime to avoid competition from conspecifics and nocturnal predators. Bears are overwhelmingly solitary and are considered to be the most asocial of all the Carnivora. The only times bears are encountered in groups are mothers with young or occasional seasonal bounties of rich food (such as salmon runs). Fights between males can occur and older individuals may have extensive scar
A scar (or scar tissue) is an area of fibrous tissue that replaces normal skin after an injury. Scars result from the biological process of wound repair in the skin, as well as in other organs, and tissues of the body. Thus, scarring is a na ...
ring, which suggests that maintaining dominance can be intense. With their acute sense of smell, bears can locate carcasses from several kilometres away. They use olfaction
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste.
In humans, it ...
to locate other foods, encounter mates, avoid rivals and recognize their cubs.
Feeding
Most bears are opportunistic omnivore
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...
s and consume more plant than animal matter, and appears to have evolved from an ancestor which was a low-protein macronutrient omnivore. They eat anything from leaves, roots, and berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
to insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s, carrion
Carrion () is the decaying flesh of dead animals, including human flesh.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
, fresh meat, and fish, and have digestive systems and teeth adapted to such a diet. At the extremes are the almost entirely herbivorous giant panda and the mostly carnivorous polar bear. However, all bears feed on any food source that becomes seasonally available. For example, Asiatic black bears in Taiwan
Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
consume large numbers of acorn
The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally
two seeds), enclosed in a tough, leathery shell, and borne ...
s when these are most common, and switch to ungulate
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
s at other times of the year.
When foraging for plants, bears choose to eat them at the stage when they are at their most nutritious and digestible, typically avoiding older grass
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es, sedge
The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
s and leaves. Hence, in more northern temperate areas, browsing and grazing is more common early in spring and later becomes more restricted. Knowing when plants are ripe for eating is a learned behavior. Berries may be foraged in bushes or at the tops of trees, and bears try to maximize the number of berries consumed versus foliage. In autumn, some bear species forage large amounts of naturally fermented
Fermentation is a metabolic process that produces chemical changes in organic substrates through the action of enzymes. In biochemistry, it is narrowly defined as the extraction of energy from carbohydrates in the absence of oxygen. In food ...
fruits, which affects their behavior. Smaller bears climb trees to obtain mast (edible reproductive parts, such as acorns). Such masts can be very important to the diets of these species, and mast failures may result in long-range movements by bears looking for alternative food sources. Brown bears, with their powerful digging abilities, commonly eat roots. The panda's diet is over 99% bamboo, of 30 different species. Its strong jaws are adapted for crushing the tough stems of these plants, though they prefer to eat the more nutritious leaves. Bromeliad
The Bromeliaceae (the bromeliads) are a family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the tropical Americas, with several species found in the American subtropics and one in tropical west Africa, ...
s can make up to 50% of the diet of the spectacled bear, which also has strong jaws to bite them open.
The sloth bear is not as specialized as polar bears and the panda, has lost several front teeth usually seen in bears, and developed a long, suctioning tongue to feed on the ant
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s, termite
Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
s, and other burrowing insects. At certain times of the year, these insects can make up 90% of their diets. Some individuals become addicted to sweets in garbage inside towns where tourism-related waste is generated throughout the year. Some species may raid the nests of wasp
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
s and bee
Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s for the honey
Honey is a sweet and viscous substance made by several bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primar ...
and immature insects, in spite of stinging from the adults. Sun bears use their long tongues to lick up both insects and honey. Fish are an important source of food for some species, and brown bears in particular gather in large numbers at salmon run
''Salmon Run'' is a 1982 video game for the Atari 8-bit family created by Bill Williams and distributed via the Atari Program Exchange. ''Salmon Run'' was the first game in Williams's career, followed by a string of successes noted for their o ...
s. Typically, a bear plunges into the water and seizes a fish with its jaws or front paws. The preferred parts to eat are the brain and eggs. Small burrowing mammals like rodent
Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s may be dug out and eaten.
The brown bear and both species of black bears sometimes take large ungulates, such as deer
Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
and bovid
The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, the ...
s, mostly the young and weak. These animals may be taken by a short rush and ambush, though hiding young may be stiffed out and pounced on. The polar bear mainly preys on seals, stalking them from the ice or breaking into their dens. They primarily eat the highly digestible blubber. Large mammalian prey is typically killed by a bite to the head or neck, or (in the case of young) simply pinned down and mauled. Predatory behavior in bears is typically taught to the young by the mother.
Bears are prolific scavenger
Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...
s and kleptoparasite
Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when fo ...
s, stealing food caches from rodents, and carcasses from other predators. For hibernating species, weight gain is important as it provides nourishment during winter dormancy. A brown bear can eat of food and gain of fat a day prior to entering its den.
Communication
Bears produce a number of vocal and non-vocal sounds. Tongue-clicking, grunting or chuffing many be made in cordial situations, such as between mothers and cubs or courting couples, while moaning, huffing, snorting or blowing air is made when an individual is stressed. Barking is produced during times of alarm, excitement or to give away the animal's position. Warning sounds include jaw-clicking and lip-popping, while teeth-chatters, bellows, growls, roars and pulsing sounds are made in aggressive encounters. Cubs may squeal, bawl, bleat or scream when in distress and make motor-like humming when comfortable or nursing.
Bears sometimes communicate with visual displays such as standing upright, which exaggerates the individual's size. The chest markings of some species may add to this intimidating display. Staring is an aggressive act and the facial markings of spectacled bears and giant pandas may help draw attention to the eyes during agonistic
An agonist is a chemical that activates a receptor to produce a biological response. Receptors are cellular proteins whose activation causes the cell to modify what it is currently doing. In contrast, an antagonist blocks the action of the agon ...
encounters. Individuals may approach each other by stiff-legged walking with the head lowered. Dominance between bears is asserted by making a frontal orientation, showing the canine teeth
In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened howeve ...
, muzzle twisting and neck stretching. A subordinate may respond with a lateral orientation, by turning away and dropping the head and by sitting or lying down.
Bears may mark territory by rubbing
A rubbing ('' frottage'') is a reproduction of the texture of a surface created by placing a piece of paper or similar material over the subject and then rubbing the paper with something to deposit marks, most commonly charcoal or pencil but a ...
against trees and other objects which may serve to spread their scent. This is usually accompanied by clawing and biting the object. Bark may be spread around to draw attention to the marking post. Pandas are known to mark objects with urine and a waxy substance from their anal glands. Polar bears leave behind their scent in their tracks which allow individuals to keep track of one another in the vast Arctic wilderness.
Reproduction and development
The mating system of bears has variously been described as a form of polygyny
Polygyny (; from Neoclassical Greek πολυγυνία (); ) is the most common and accepted form of polygamy around the world, entailing the marriage of a man with several women.
Incidence
Polygyny is more widespread in Africa than in any o ...
, promiscuity
Promiscuity is the practice of engaging in sexual activity frequently with different Sexual partner, partners or being indiscriminate in the choice of sexual partners. The term can carry a moral judgment. A common example of behavior viewed as pro ...
and serial monogamy
Monogamy ( ) is a form of dyadic relationship in which an individual has only one partner during their lifetime. Alternately, only one partner at any one time (serial monogamy) — as compared to the various forms of non-monogamy (e.g., polyga ...
. During the breeding season, males take notice of females in their vicinity and females become more tolerant of males. A male bear may visit a female continuously over a period of several days or weeks, depending on the species, to test her reproductive state. During this time period, males try to prevent rivals from interacting with their mate. Courtship may be brief, although in some Asian species, courting pairs may engage in wrestling, hugging, mock fighting and vocalizing. Ovulation is induced by mating, which can last up to 30 minutes depending on the species.
Gestation
Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
typically lasts 6–9 months, including delayed implantation Embryonic diapause (from late 19th century English: dia- ‘through’ + pause- 'delay') (aka delayed implantation in mammals) is an evolutionary reproductive strategy used by several animal species across a number of kingdoms, including approximate ...
, and litter size numbers up to four cubs. Giant pandas may give birth to twins but they can only suckle one young and the other is left to die. In northern living species, birth takes place during winter dormancy. Cubs are born blind and helpless with at most a thin layer of hair, relying on their mother for warmth. The milk of the female bear is rich in fat and antibodies and cubs may suckle for up to a year after they are born. By 2–3 months, cubs can follow their mother outside the den. They usually follow her on foot, but sloth bear cubs may ride on their mother's back. Male bears play no role in raising young. Infanticide
Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of reso ...
, where an adult male kills the cubs of another, has been recorded in polar bears, brown bears and American black bears but not in other species. Males kill young to bring the female into estrus
The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
. Cubs may flee and the mother defends them even at the cost of her life.
In some species, offspring may become independent around the next spring, though some may stay until the female successfully mates again. Bears reach sexual maturity
Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definitio ...
shortly after they disperse; at around 3–6 years depending on the species. Male Alaskan brown bears and polar bears may continue to grow until they are 11 years old. Lifespan may also vary between species. The brown bear can live an average of 25 years.
Hibernation
Bears of northern regions, including the American black bear
The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
and the grizzly bear
The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America.
In addition to the mainland grizzly (''Ursus arctos horri ...
, hibernate in the winter. During hibernation, the bear's metabolism slows down, its body temperature decreases slightly, and its heart rate slows from a normal value of 55 to just 9 beats per minute. Bears normally do not wake during their hibernation, and can go the entire period without eating, drinking, urinating, or defecating. A fecal plug A fecal plug (occasionally known as a ''tappen'') is a large mass of hardened feces produced by a bear during its winter hibernation. The plug forms in the colon, and consists of a variety of materials ingested by the bear during and immediately be ...
is formed in the colon, and is expelled when the bear wakes in the spring. If they have stored enough body fat, their muscles remain in good condition, and their protein maintenance requirements are met from recycling waste urea
Urea, also known as carbamide, is an organic compound with chemical formula . This amide has two amino groups (–) joined by a carbonyl functional group (–C(=O)–). It is thus the simplest amide of carbamic acid.
Urea serves an important r ...
. Female bears give birth during the hibernation period, and are roused when doing so.
Mortality
Bears do not have many predators. The most important are humans, and as they started cultivating crops, they increasingly came in conflict with the bears that raided them. Since the invention of firearms, people have been able to kill bears with greater ease. Felid
Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the dom ...
s like the tiger
The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...
may also prey on bears, particularly cubs, which may also be threatened by canids.[
Bears are parasitized by eighty species of parasites, including single-celled protozoans and gastro-intestinal worms, and nematodes and flukes in their heart, liver, lungs and bloodstream. Externally they have ticks, fleas and lice. A study of American black bears found seventeen species of endoparasite including the protozoan '']Sarcocystis
''Sarcocystis'' is a genus of protozoan parasites, with many species infecting mammals, reptiles and birds. Its name is dervived from Greek ''sarx'' = flesh and ''kystis'' = bladder.
The lifecycle of a typical member of this genus involves two ...
'', the parasitic worm ''Diphyllobothrium mansonoides
''Diphyllobothrium mansonoides'' (also known as ''Spirometra mansonoides'') is a species of tapeworm (cestodes) that is endemic to North America. Infection with ''D. mansonoides'' in humans can result in sparganosis. Justus F. Mueller first repor ...
'', and the nematodes ''Dirofilaria immitis
''Dirofilaria immitis'', also known as heartworm or dog heartworm, is a parasitic roundworm that is a type of filarial worm, a small thread-like worm, that causes dirofilariasis. It is spread from host to host through the bites of mosquitoes ...
'', ''Capillaria aerophila
''Capillaria aerophila'' is a nematode parasite found in the respiratory tract of foxes, dogs, and various other carnivorous mammals. A few cases of human infestation have also been reported. Though it is sometimes called a "lungworm", this ter ...
'', ''Physaloptera
Systematics
''Physaloptera'' is a genus of parasitic nematodes in the family Physalopteridae. Species include:
*''Physaloptera dilatata''Muniz-Pereira et al., 2009, p. 11
*''Physaloptera dispar''
*''Physaloptera hispida''Kinsella, 1974, p ...
'' sp., ''Strongyloides
''Strongyloides'' (from Greek ''strongylos'', round, + ''eidos'', resemblance), anguillula, or threadworm is a genus of small nematode parasites, belonging to the family Strongylidae, commonly found in the small intestine of mammals (particularl ...
'' sp. and others. Of these, ''D. mansonoides'' and adult ''C. aerophila'' were causing pathological symptoms. By contrast, polar bears have few parasites; many parasitic species need a secondary, usually terrestrial, host, and the polar bear's life style is such that few alternative hosts exist in their environment. The protozoan ''Toxoplasma gondii
''Toxoplasma gondii'' () is an obligate intracellular parasitic protozoan (specifically an apicomplexan) that causes toxoplasmosis. Found worldwide, ''T. gondii'' is capable of infecting virtually all warm-blooded animals, but felids, such as d ...
'' has been found in polar bears, and the nematode ''Trichinella nativa
''Trichinella nativa'' is a nematode worm, one of the species of the genus ''Trichinella'', found in arctic and subarctic regions.
Biology
It is highly pathogenic and has a high resistance to freezing. It is encapsulated, and infects a wide va ...
'' can cause a serious infection and decline in older polar bears. Bears in North America are sometimes infected by a ''Morbillivirus
''Morbillivirus'' is a genus of viruses in the order ''Mononegavirales'', in the family ''Paramyxoviridae''. Humans, dogs, cats, cattle, seals, and cetaceans serve as natural hosts. This genus includes seven species. Diseases in humans associate ...
'' similar to the canine distemper virus
Canine may refer to:
Zoology and anatomy
* a dog-like Canid animal in the subfamily Caninae
** ''Canis'', a genus including dogs, wolves, coyotes, and jackals
** Dog, the domestic dog
* Canine tooth, in mammalian oral anatomy
People with the surn ...
. They are susceptible to infectious canine hepatitis
Infectious canine hepatitis (ICH) is an acute liver infection in dogs caused by ''Canine mastadenovirus A'', formerly called ''Canine adenovirus 1'' (CAV-1). CAV-1 also causes disease in wolves, coyotes, and bears, and encephalitis in foxes. The ...
(CAV-1), with free-living black bears dying rapidly of encephalitis and hepatitis.
Relationship with humans
Conservation
In modern times, bears have come under pressure through encroachment on their habitats and illegal trade in bear parts, including the Asian bile bear
Bile bears, sometimes called battery bears, are bears kept in captivity to harvest their bile, a digestive fluid produced by the liver and stored in the gallbladder, which is used by some traditional Asian medicine practitioners. It is estimated t ...
market, though hunting is now banned, largely replaced by farming. The IUCN
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
lists six bear species as vulnerable; even the two least concern
A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
species, the brown bear and the American black bear,[ are at risk of ]extirpation
Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
in certain areas. In general these two species inhabit remote areas with little interaction with humans, and the main non-natural causes of mortality are hunting, trapping, road-kill and depredation.
Laws have been passed in many areas of the world to protect bears from habitat destruction
Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
. Public perception of bears is often positive, as people identify with bears due to their omnivorous diets, their ability to stand on two legs, and their symbolic importance. Support for bear protection is widespread, at least in more affluent societies. The giant panda has become a worldwide symbol of conservation. The Sichuan Giant Panda Sanctuaries
The Sichuan Giant Panda Sanctuaries () located in southwest Sichuan province of China, is the home to more than 30% of the world's giant pandas and is among the most important sites for the captive breeding of these pandas. It covers 9245 km2 ...
, which are home to around 30% of the wild panda population, gained a UNESCO World Heritage Site
A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
designation in 2006. Where bears raid crops or attack livestock, they may come into conflict with humans. In poorer rural regions, attitudes may be more shaped by the dangers posed by bears, and the economic costs they cause to farmers and ranchers.
Attacks
Several bear species are dangerous to humans, especially in areas where they have become used to people; elsewhere, they generally avoid humans. Injuries caused by bears are rare, but are widely reported. Bears may attack humans in response to being startled, in defense of young or food, or even for predatory reasons.
Entertainment, hunting, food and folk medicine
Bears in captivity have for centuries been used for entertainment. They have been trained to dance
Dance is a performing art form consisting of sequences of movement, either improvised or purposefully selected. This movement has aesthetic and often symbolic value. Dance can be categorized and described by its choreography, by its repertoir ...
, and were kept for baiting in Europe at least since the 16th century. There were five bear-baiting gardens in Southwark
Southwark ( ) is a district of Central London situated on the south bank of the River Thames, forming the north-western part of the wider modern London Borough of Southwark. The district, which is the oldest part of South London, developed ...
, London, at that time; archaeological remains of three of these have survived. Across Europe, nomad
A nomad is a member of a community without fixed habitation who regularly moves to and from the same areas. Such groups include hunter-gatherers, pastoral nomads (owning livestock), tinkers and trader nomads. In the twentieth century, the popu ...
ic Romani
Romani may refer to:
Ethnicities
* Romani people, an ethnic group of Northern Indian origin, living dispersed in Europe, the Americas and Asia
** Romani genocide, under Nazi rule
* Romani language, any of several Indo-Aryan languages of the Roma ...
bear handlers called Ursari
The Ursari (generally read as " bear leaders" or "bear handlers"; from the ro, urs, meaning "bear"; singular: ''ursar''; Bulgarian: урсари, ''ursari'') or Richinara are the traditionally nomadic occupational group of animal trainers among ...
lived by busking with their bears from the 12th century.
Bears have been hunted for sport, food, and folk medicine
Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
. Their meat is dark and stringy, like a tough cut of beef. In Cantonese cuisine
Cantonese or Guangdong cuisine, also known as Yue cuisine ( or ) is the cuisine of Guangdong province of China, particularly the provincial capital Guangzhou, and the surrounding regions in the Pearl River Delta including Hong Kong and Maca ...
, bear paws are considered a delicacy. Bear meat should be cooked thoroughly, as it can be infected with the parasite ''Trichinella spiralis
''Trichinella spiralis'' is a viviparous nematode parasite, occurring in rodents, pigs, bears, hyenas and humans, and is responsible for the disease trichinosis. It is sometimes referred to as the "pork worm" due to it being typically encounte ...
''.
The peoples of eastern Asia use bears' body parts and secretions (notably their gallbladders and bile) as part of traditional Chinese medicine
Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of action ...
. More than 12,000 bears are thought to be kept on farms in China, Vietnam, and South Korea for the production of bile. Trade in bear products is prohibited under CITES
CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
, but bear bile has been detected in shampoos, wine and herbal medicines sold in Canada, the United States and Australia.
File:The dancing bear by William Frederick Witherington.jpg, ''The Dancing Bear'' by William Frederick Witherington
William Frederick Witherington (26 May 1785 – 10 April 1865) was an English painter and academic. Born in London, he entered the Royal Academy Schools in 1805. Except for one year he exhibited annually at the Royal Academy from 1811 until his ...
, 1822
File:Theodor Aman - Ursarul.jpg, A nomadic ''ursar
The Ursari (generally read as " bear leaders" or "bear handlers"; from the ro, urs, meaning "bear"; singular: ''ursar''; Bulgarian: урсари, ''ursari'') or Richinara are the traditionally nomadic occupational group of animal trainers among ...
'', a Romani
Romani may refer to:
Ethnicities
* Romani people, an ethnic group of Northern Indian origin, living dispersed in Europe, the Americas and Asia
** Romani genocide, under Nazi rule
* Romani language, any of several Indo-Aryan languages of the Roma ...
bear-busker. Drawing by Theodor Aman
Theodor Aman (20 March 1831 – 19 August 1891) was a Romanian painter, engraver and art professor. He mostly produced genre and history scenes.
Biography
His father was a cavalry commander from Craiova but he was born in Câmpulung, where his f ...
, 1888
Cultural depictions
Bears have been popular subjects in art, literature, folklore and mythology. The image of the mother bear was prevalent throughout societies in North America and Eurasia, based on the female's devotion and protection of her cubs. In many Native American cultures, the bear is a symbol of rebirth because of its hibernation and re-emergence. A widespread belief among cultures of North America and northern Asia associated bears with shaman
Shamanism is a religious practice that involves a practitioner (shaman) interacting with what they believe to be a spirit world through altered states of consciousness, such as trance. The goal of this is usually to direct spirits or spiritu ...
; this may be based on the solitary nature of both. Bears have thus been thought to predict the future and shaman were believed to have been capable of transforming into bears.
There is evidence of prehistoric bear worship
Bear worship (also known as the bear cult or arctolatry) is the religious practice of the worshipping of bears found in many North Eurasian ethnic religions such as among the Sami, Nivkh, Ainu, Basques, Germanic peoples, Slavs and Finns. There a ...
, though this is disputed by archaeologists. It is possible that bear worship existed in early Chinese and Ainu culture
Ainu culture is the culture of the Ainu people, from around the 13th century (late Kamakura period) to the present. Today, most Ainu people live a life superficially similar to that of mainstream Japanese people, partly due to cultural assimilat ...
s. The prehistoric Finns
Finns or Finnish people ( fi, suomalaiset, ) are a Baltic Finnic ethnic group native to Finland.
Finns are traditionally divided into smaller regional groups that span several countries adjacent to Finland, both those who are native to these ...
, Siberian peoples
Siberia, including the Russian Far East, is a vast region spanning the North Asia, northern part of the Asian continent, and forming the Asiatic portion of Russia. As a result of the Russian conquest of Siberia (17th to 19th centuries) and of t ...
and more recently Koreans
Koreans ( South Korean: , , North Korean: , ; see names of Korea) are an East Asian ethnic group native to the Korean Peninsula.
Koreans mainly live in the two Korean nation states: North Korea and South Korea (collectively and simply refe ...
considered the bear as the spirit of their forefathers. Artio
Artio (''Dea Artio'' in the Gallo-Roman religion) is a Celtic bear goddess. Evidence of her worship has notably been found at Bern in Switzerland. Her name is derived from the Gaulish word for 'bear', ''artos''.
Name
The Gaulish theonym ''Artiō' ...
(''Dea Artio'' in the Gallo-Roman religion
Gallo-Roman religion is a fusion of the traditional religious practices of the Gauls, who were originally Celtic speakers, and the Roman and Hellenistic religions introduced to the region under Roman Imperial rule. It was the result of selective ...
) was a Celtic
Celtic, Celtics or Keltic may refer to:
Language and ethnicity
*pertaining to Celts, a collection of Indo-European peoples in Europe and Anatolia
**Celts (modern)
*Celtic languages
**Proto-Celtic language
* Celtic music
*Celtic nations
Sports Fo ...
bear goddess. Evidence of her worship has notably been found at Bern
german: Berner(in)french: Bernois(e) it, bernese
, neighboring_municipalities = Bremgarten bei Bern, Frauenkappelen, Ittigen, Kirchlindach, Köniz, Mühleberg, Muri bei Bern, Neuenegg, Ostermundigen, Wohlen bei Bern, Zollikofen
, website ...
, itself named for the bear. Her name is derived from the Celtic word for "bear", ''artos''. In ancient Greece
Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cult ...
, the archaic cult of Artemis
In ancient Greek mythology and religion, Artemis (; grc-gre, Ἄρτεμις) is the goddess of the hunt, the wilderness, wild animals, nature, vegetation, childbirth, care of children, and chastity. She was heavily identified wit ...
in bear form survived into Classical times at Brauron
Brauron (; grc, Βραυρών) was one of the twelve cities of ancient Attica, but never mentioned as a ''deme'', though it continued to exist down to the latest times. It was situated on or near the eastern coast of Attica, between Steiria and H ...
, where young Athenian girls passed an initiation rite as ''arktoi'' "she bears".
The constellations of Ursa Major and Ursa Minor
Ursa Minor (Latin: 'Lesser Bear', contrasting with Ursa Major), also known as the Little Bear, is a constellation located in the far northern sky. As with the Great Bear, the tail of the Little Bear may also be seen as the handle of a ladle, h ...
, the great and little bears, are named for their supposed resemblance to bears, from the time of Ptolemy
Claudius Ptolemy (; grc-gre, Πτολεμαῖος, ; la, Claudius Ptolemaeus; AD) was a mathematician, astronomer, astrologer, geographer, and music theorist, who wrote about a dozen scientific treatises, three of which were of importanc ...
.[ The nearby star ]Arcturus
, - bgcolor="#FFFAFA"
, Note (category: variability): , , H and K emission vary.
Arcturus is the brightest star in the northern constellation of Boötes. With an apparent visual magnitude of −0.05, it is the third-brightest of th ...
means "guardian of the bear", as if it were watching the two constellations. Ursa Major has been associated with a bear for as much as 13,000 years since Paleolithic
The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...
times, in the widespread Cosmic Hunt
The Cosmic Hunt is an old and widely distributed family of cognate myths. They are stories about a large animal that is pursued by hunters, is wounded, and is transformed into a constellation. Variants of the Cosmic Hunt are common in cultures of ...
myths. These are found on both sides of the Bering land bridge
Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of ...
, which was lost to the sea some 11,000 years ago.
Bears are popular in children's stories, including Winnie the Pooh
Winnie-the-Pooh, also called Pooh Bear and Pooh, is a fictional Anthropomorphism, anthropomorphic teddy bear created by English author A. A. Milne and English illustrator E. H. Shepard.
The first collection of stories about the character w ...
, Paddington Bear
Paddington Bear is a fictional character in children's literature. He first appeared on 13 October 1958 in the children's book ''A Bear Called Paddington'' and has been featured in more than twenty books written by British author Michael Bond, a ...
, Gentle Ben
Gentle Ben is a bear character created by author Walt Morey and first introduced in a 1965 children's novel, ''Gentle Ben''. The original novel told the story of the friendship between a large male bear named Ben and a boy named Mark. The story p ...
and "The Brown Bear of Norway
The Brown Bear of Norway is an Irish fairy tale collected by Patrick Kennedy which appeared in his ''Legendary Fictions of the Irish Celts'' (1866). It was later included by Andrew Lang in his anthology '' The Lilac Fairy Book'' (1910), though La ...
". An early version of "Goldilocks and the Three Bears
"Goldilocks and the Three Bears" (originally titled "The Story of the Three Bears") is a 19th-century English fairy tale of which three versions exist. The original version of the tale tells of an obscene old woman who enters the forest home ...
", was published as "The Three Bears" in 1837 by Robert Southey
Robert Southey ( or ; 12 August 1774 – 21 March 1843) was an English poet of the Romantic school, and Poet Laureate from 1813 until his death. Like the other Lake Poets, William Wordsworth and Samuel Taylor Coleridge, Southey began as a ra ...
, many times retold, and illustrated in 1918 by Arthur Rackham
Arthur Rackham (19 September 1867 – 6 September 1939) was an English book illustrator. He is recognised as one of the leading figures during the Golden Age of British book illustration. His work is noted for its robust pen and ink drawings, ...
. The Hanna-Barbera
Hanna-Barbera Cartoons, Inc. ( ) was an American animation studio and production company which was active from 1957 to 2001. It was founded on July 7, 1957, by William Hanna and Joseph Barbera following the decision of Metro-Goldwyn-Mayer to c ...
character Yogi Bear
Yogi Bear is an anthropomorphic animal character who has appeared in numerous comic books, animated television shows and films. He made his debut in 1958 as a supporting character in ''The Huckleberry Hound Show''.
Yogi Bear was the first ...
has appeared in numerous comic books, animated television show
An animated series is a set of animated works with a common series title, usually related to one another. These episodes should typically share the same main characters, some different secondary characters and a basic theme. Series can have either ...
s and films. The Care Bears
Care Bears are multi-colored bears, originally painted in 1981 by artist Elena Kucharik to be used on greeting cards from American Greetings. In 1983, the characters were turned into plush teddy bears.
The characters headlined their own televisi ...
began as greeting cards in 1982, and were featured as toys, on clothing and in film. Around the world, many children—and some adults—have teddy bear
A teddy bear is a stuffed toy in the form of a bear. Developed apparently simultaneously by toymakers Morris Michtom in the U.S. and Richard Steiff under his aunt Margarete Steiff's company in Germany in the early 20th century, the teddy bear, ...
s, stuffed toy
A stuffed toy is a toy doll with an outer fabric sewn from a textile and stuffed with flexible material. They are known by many names, such as plush toys, plushies, stuffed animals, and stuffies; in Britain and Australia, they may also be cal ...
s in the form of bears, named after the American statesman Theodore Roosevelt
Theodore Roosevelt Jr. ( ; October 27, 1858 – January 6, 1919), often referred to as Teddy or by his initials, T. R., was an American politician, statesman, soldier, conservationist, naturalist, historian, and writer who served as the 26t ...
when in 1902 he had refused to shoot an American black bear tied to a tree.
Bears, like other animals, may symbolize nations. The Russian Bear has been a common national personification
A national personification is an anthropomorphic personification of a state or the people(s) it inhabits. It may appear in political cartoons and propaganda.
Some early personifications in the Western world tended to be national manifestations ...
for Russia from the 16th century onward. Smokey Bear
Smokey Bear is an American campaign and advertising icon of the U.S. Forest Service. In the Wildfire Prevention Campaign, which is the longest-running public service announcement campaign in United States history, the Ad Council, the United St ...
has become a part of American culture since his introduction in 1944, with his message "Only you can prevent forest fires".
File:The Three Bears - Project Gutenberg eText 17034.jpg, "The Three Bears", Arthur Rackham
Arthur Rackham (19 September 1867 – 6 September 1939) was an English book illustrator. He is recognised as one of the leading figures during the Golden Age of British book illustration. His work is noted for its robust pen and ink drawings, ...
's illustration to ''English Fairy Tales'', by Flora Annie Steel
Flora Annie Steel (2 April 1847 – 12 April 1929) was a writer who lived in British India for 22 years. She was noted especially for books set in the Indian sub-continent or connected with it. Her novel ''On the Face of the Waters'' (1896) desc ...
, 1918
File:Sidney Hall - Urania's Mirror - Ursa Major.jpg, The constellation
A constellation is an area on the celestial sphere in which a group of visible stars forms Asterism (astronomy), a perceived pattern or outline, typically representing an animal, mythological subject, or inanimate object.
The origins of the e ...
of Ursa Major
Ursa Major (; also known as the Great Bear) is a constellation in the northern sky, whose associated mythology likely dates back into prehistory. Its Latin name means "greater (or larger) bear," referring to and contrasting it with nearby Ursa ...
as depicted in Urania's Mirror
''Urania's Mirror; or, a view of the Heavens'' is a set of 32 astronomical star chart cards, first published in November 1824. They are illustrations based on Alexander Jamieson's ''A Celestial Atlas'', but the addition of holes punched in them ...
, c. 1825
Organizations
The International Association for Bear Research & Management, also known as the International Bear Association The International Association for Bear Research and Management (IBA), sometimes shortened to International Bear Association, is a professional organization for biologists, wildlife managers and others which focuses on wildlife conservation of the ei ...
, and the Bear Specialist Group
Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the North ...
of the Species Survival Commission, a part of the International Union for Conservation of Nature
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
focus on the natural history, management, and conservation of bears. Bear Trust International works for wild bears and other wildlife through four core program initiatives, namely Conservation Education, Wild Bear Research, Wild Bear Management, and Habitat Conservation.
Specialty organizations for each of the eight species of bears worldwide include:
* Vital Ground, for the brown bear
* Moon Bears, for the Asiatic black bear
* Black Bear Conservation Coalition, for the North American black bear
* Polar Bears International, for the polar bear
* Bornean Sun Bear Conservation Centre, for the sun bear
* Wildlife SOS, for the sloth bear
* Andean Bear Conservation Project, for the Andean bear
* Chengdu Research Base of Giant Panda Breeding, for the giant panda
See also
* List of fictional bears
This is a list of fictional bears that appear in video games, film, television, animation, comics and literature. This also includes pandas, but not the unrelated red panda species. The list is limited to notable, named characters. This list is a ...
* List of individual bears
The following is a list of individual bears which garnered national or worldwide attention:
Actors
* Bart the Bear, a male Alaskan Kodiak bear, played the leading role in the 1988 wilderness drama, ''The Bear (1988 film), The Bear''. Between 1980 ...
Notes
References
Citations
General and cited references
*
Further reading
*
*
*
External links
The Bears Project – Information, reports and images of European brown bears and other living species
The Bear Book and Curriculum Guide – a compilation of stories about all eight species of bears worldwide, including STEM lessons rooted in bear research, ecology, and conservation
{{Authority control
Articles containing video clips
*
Extant Eocene first appearances
Predators
Taxa named by Gotthelf Fischer von Waldheim