Barsboldia Vertebrae
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''Barsboldia'' (meaning "of Barsbold", a well-known Mongolian paleontologist) is a genus of large
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
dinosaur from the
early Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval fro ...
Nemegt Formation of Ömnogöv', Mongolia. It is known from a partial vertebral column, partial
pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
, and some ribs.


Discovery

In 1970, a Polish-Mongolian expedition near the Nemegt found the skeleton of an ornithopod and first assigned it to ''
Saurolophus angustirostris ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 million ...
''. However, two Polish paleontologists, Teresa Maryańska and
Halszka Osmólska Halszka Osmólska (September 15, 1930 – March 31, 2008) was a Polish paleontologist who had specialized in Mongolian dinosaurs. Biography She was born in 1930 in Poznań. In 1949, she began to study biology at Faculty of Biology and Earth Scie ...
, came to the conclusion that it was a
lambeosaurine Lambeosaurinae is a group of crested hadrosaurid dinosaurs. Classification Lambeosaurines have been traditionally split into the tribes or clades Parasaurolophini ('' Parasaurolophus'', ''Charonosaurus'', others (?).) and Lambeosaurini (''Cor ...
that had to represent a separate species. They named and described ''Barsboldia sicinskii'' based on the holotype specimen ZPAL MgD-1/110. The genus name honors the Mongolian paleontologist Rinchen Barsbold, while the species name honors Wojciech Siciński, the technician at the Warsaw Paleobiological Institute who prepared the skeleton. The holotype specimen, found in a layer of the Nemegt Formation dating to the early Maastrichtian, consisted of a partial skeleton consisting of nine back vertebrae, nine hip vertebrae, fifteen tail vertebrae, a left
ilium Ilium or Ileum may refer to: Places and jurisdictions * Ilion (Asia Minor), former name of Troy * Ilium (Epirus), an ancient city in Epirus, Greece * Ilium, ancient name of Cestria (Epirus), an ancient city in Epirus, Greece * Ilium Building, a ...
, parts of the left and right pubis, several ribs, and a few fragments of the hind limbs, with the backbone largely articulated. The anterior and posteriormost portions of the skeleton were lost due to erosion.


Description

Barsboldia was a large hadrosaur, previously measured in length and in body mass. In 2011, the tibial length was measured at , rivalling that of ''
Shantungosaurus ''Shantungosaurus'' (meaning "''Shandong Lizard''") is a genus of very large saurolophine hadrosaurid dinosaur found in the Late Cretaceous Wangshi Group of the Shandong Peninsula in China, containing a single species, ''Shantungosaurus giganteu ...
'' measured at and that of ''
Magnapaulia ''Magnapaulia'' is a genus of herbivorous lambeosaurine hadrosaurid dinosaurs known from the Latest Cretaceous Baja California, of northwestern Mexico. It contains a single species, ''Magnapaulia laticaudus''. ''Magnapaulia'' was first described ...
'' measured at ; this indicates that ''Barsboldia'' could have possibly reached within the range of in total body length. The most distinctive features of ''Barsboldia'' are found in the
neural spines The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
. These are very tall, particularly over the hips, and were described as second only to those of ''
Hypacrosaurus altispinus ''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high because it was almost but not quite as large as ''Tyrannosaurus'') was a genus of duckbill dinosaur similar in appearance to ''Co ...
'' and the tips of those found in the first few vertebrae of the tail are club-shaped, possibly a sign of old age.


Phylogeny

Maryańska and Osmólska described their new genus as a
lambeosaurine Lambeosaurinae is a group of crested hadrosaurid dinosaurs. Classification Lambeosaurines have been traditionally split into the tribes or clades Parasaurolophini ('' Parasaurolophus'', ''Charonosaurus'', others (?).) and Lambeosaurini (''Cor ...
(or hollow-crested duckbill), the first from the Nemegt Formation, although it lacked a skull. However, the sacrum has a keel along the bottom, a possible lambeosaurine feature, and the bones closely resemble those of ''Hypacrosaurus''. With only one partial skeleton known, and no skull, the genus has been considered dubious or a possible lambeosaurine of uncertain placement. A newer study published in 2011 suggests that ''Barsboldia'' is actually a valid
saurolophine Saurolophinae is a subfamily (biology), subfamily of hadrosaurid dinosaurs. It has since the mid-20th century generally been called the Hadrosaurinae, a group of largely non-crested hadrosaurs related to the crested sub-family Lambeosaurinae. How ...
. The following cladogram was recovered in the 2011 phylogenetic analysis of Hadrosauroidea by Prieto-Márquez (the relationships within Lambeosaurinae and between basal hadrosauroids aren't shown).


Paleobiology

As a hadrosaurid, ''Barsboldia'' would have been a bipedal/ quadrupedal
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, eating plants with a sophisticated skull that permitted a grinding motion analogous to chewing, and was furnished with hundreds of continually-replaced teeth. If it was a lambeosaurine, it would have had a hollow crest formed out of expanded skull bones containing the nasal passages, with a function relating to identification by sight and sound.


See also

* Timeline of hadrosaur research


References

{{Taxonbar, from1=Q18606529, from2=Q264109 Saurolophines Late Cretaceous dinosaurs of Asia Maastrichtian life Nemegt fauna Fossil taxa described in 1981 Taxa named by Teresa Maryańska Taxa named by Halszka Osmólska Ornithischian genera