Barred Forest Falcon
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The barred forest falcon (''Micrastur ruficollis'') is a species of
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in subfamily
Herpetotherinae Herpetotherinae is a subfamily of falconid birds of prey that includes eight species in two genera '' Herpetotheres'' (laughing falcons) and '' Micrastur'' (forest falcons). Both genera are found in South America and the subfamily is basal to th ...
of family
Falconidae The falcons and caracaras are around 60 species of diurnal birds of prey that make up the family Falconidae (representing all extant species in the order Falconiformes). The family is divided into three subfamilies, Herpetotherinae, which inclu ...
, the
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
s and caracaras. It occurs from southern Mexico south through most of Central America and in every mainland South American country except Chile and Uruguay.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. 30 January 2023. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved January 30, 2023


Taxonomy and systematics

The barred forest falcon has these six subspecies: *''M. r. guerilla'' Cassin, 1849 *''M. r. interstes''
Bangs Bang or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Lorestan, a village in I ...
, 1907
*''M. r. zonothorax'' (
Cabanis Cabanis is the surname of: *George Cabanis (1815-1892), American politician *Jean Cabanis (1816–1906), German ornithologist *José Cabanis José Cabanis (2 March 1922 – 6 October 2000) was a French novelist, essayist, historian and magistrate ...
, 1866)
*''M. r. concentricus'' (
Lesson A lesson or class is a structured period of time where learning is intended to occur. It involves one or more students (also called pupils or learners in some circumstances) being taught by a teacher or instructor. A lesson may be either one ...
, 1830)
*''M. r. ruficollis'' (
Vieillot Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist. Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collect ...
, 1817)
*''M. r. olrogi'' Amadon, 1964 What is now the lined forest falcon (''M. gilvicollis'') was previously included as a seventh subspecies.


Description

The barred forest falcon is long. Males weigh and females . Adults of most
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are typically dark slate gray above; the tail is tipped with white and has three to six narrow white bars. The throat is pale gray, shading to the darker slate of the crown. The rest of the under parts, including the under-wing coverts, are white, finely and clearly barred with black or dark gray. The upper breast is a darker gray. The primary
remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
are dark brownish-gray with off-white bars on the inner webs. Subspecies, ''M. r. zonothorax'' is polymorphic (at least in the northern part of its range), and also occurs in a brown
morph Morph may refer to: Biology * Morph (zoology), a visual or behavioral difference between organisms of distinct populations in a species * Muller's morphs, a classification scheme for genetic mutations * "-morph", a suffix commonly used in tax ...
, where most of the upperparts, head, and chest are brown or
rufous Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
instead of gray. The
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''M. r. ruficollis'' appears to only occur in the rufous-brown morphotype, as also suggested by its scientific name, ''M. r. ruficollis''.Bierregaard, R. O. and G. M. Kirwan (2020). Barred Forest-Falcon (''Micrastur ruficollis''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.baffal1.01 retrieved February 11, 2023 The eyes are
cream Cream is a dairy product composed of the higher-fat layer skimmed from the top of milk before homogenization. In un-homogenized milk, the fat, which is less dense, eventually rises to the top. In the industrial production of cream, this process ...
to light orange brown; the
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
black, becoming yellow at base of the lower mandible; the
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food, ...
, lores, and orbit are yellow, and the legs are orange-yellow.


Distribution and habitat

The subspecies of the barred forest falcon are found thus: *''M. r. guerilla'', from
San Luis Potosí San Luis Potosí (), officially the Free and Sovereign State of San Luis Potosí ( es, Estado Libre y Soberano de San Luis Potosí), is one of the 32 states which compose the Federal Entities of Mexico. It is divided in 58 municipalities and i ...
in central Mexico south to Nicaragua, but excluding the
Yucatán Peninsula The Yucatán Peninsula (, also , ; es, Península de Yucatán ) is a large peninsula in southeastern Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north ...
and El Salvador *''M. r. interstes'', from Costa Rica south through Panama, western Colombia, and western Ecuador into extreme northwestern Peru *''M. r. zonothorax'', the
Venezuelan Coastal Range The Venezuelan Coastal Range ( es, Cordillera de la Costa or ), also known as Venezuelan Caribbean Mountain System ( es, Sistema Montañoso Caribe) is a mountain range system and one of the eight natural regions of Venezuela, that runs along the c ...
, the
Serranía del Perijá The Serranía del Perijá, Cordillera de Perijá or Sierra de Perijá is a mountain range, an extension of the eastern Andean branch ( Cordillera Oriental), in northern South America, between Colombia and Venezuela Venezuela (; ), offici ...
on the Venezuela-Colombia border, Colombia's
Sierra Nevada de Santa Marta The Sierra Nevada de Santa Marta (English: ''Snow-Covered Mountain Range of Saint Martha'') is an isolated mountain range in northern Colombia, separate from the Andes range that runs through the north of the country. Reaching an elevation of ...
and Cordillera Oriental, and south in the foothills along the east side of the Andes through Ecuador and Peru into Boliva *''M. r. concentricus'', southern Venezuela,
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * ...
, and the
Amazon Basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
of Brazil *''M. r. ruficollis'', Brazil south of Amazonia, Paraguay, and
Formosa Taiwan, officially the Republic of China (ROC), is an island country located in East Asia. The main island of Taiwan, formerly known in the Western political circles, press and literature as Formosa, makes up 99% of the land area of the territorie ...
, Chaco, and
Misiones Misiones (, ''Missions'') is one of the 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamia region. It is surrounded by Paraguay to the northwest, Brazil to the north, east and south, and Corrientes P ...
provinces of north-central and northeastern Argentina *''M. r. olrogi'',
Jujuy San Salvador de Jujuy (), commonly known as Jujuy and locally often referred to as San Salvador, is the capital and largest city of Jujuy Province in Argentine Northwest, northwest Argentina. Also, it is the seat of the Doctor Manuel Belgrano Depa ...
,
Salta Salta () is the capital and largest city in the Argentine province of the same name. With a population of 618,375 according to the 2010 census, it is also the 7th most-populous city in Argentina. The city serves as the cultural and economic ce ...
and Tucumán provinces of northwestern Argentina Barred forest falcons mainly use mature upland forest. In Central America, the species is generally restricted to mature
tropical forest Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds. Some tropical fores ...
. In South America, however, the barred forest falcon lives in other kinds of forest and
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
, even relatively arid ones. For example, in the
Amazon biome The Amazon biome ( pt, Bioma Amazônia) contains the Amazon rainforest, an area of tropical rainforest, and other ecoregions that cover most of the Amazon basin and some adjacent areas to the north and east. The biome contains blackwater and white ...
it occurs most often in
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has re-grown after a timber harvest or clearing for agriculture, until a long enough period has passed so that the effects of the disturbance are no longer evident. ...
,
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, tidal swamp forest,
semideciduous Semi-deciduous or semi-evergreen is a botanical term which refers to plants that lose their foliage for a very short period, when old leaves fall off and new foliage growth is starting. This phenomenon occurs in tropical and sub-tropical woody spe ...
forest, and forest edges. In
Acre The acre is a unit of land area used in the imperial Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imp ...
, Brazil, the barred forest falcon is reported to prefer disturbed forest types, both natural secondary and man-made, including
bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
and more open seasonally drier forest on rocky outcrops, but generally it is a bird that avoids habitat where human influence is too pronounced, and requires primary or mature secondary forest to persist in any location.Laverde-R., Oscar; Stiles, F. Gary & Múnera-R., Claudia (2005): Nuevos registros e inventario de la avifauna de la Serranía de las Quinchas, un área importante para la conservación de las aves (AICA) en Colombia ew records and updated inventory of the avifauna of the Serranía de las Quinchas, an important bird area (IBA) in Colombia ''Caldasia'' 27(2): 247-265 panish with English abstractbr>PDF fulltext
/ref> The barred forest falcon is rare on the eastern slope of Colombia's Cordillera Oriental, where it was recorded in
primary forest An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological featur ...
and old secondary forest, in a narrow elevational band between Second-growth forest in these mountains is dominated by trees like
Melastomaceae Melastomataceae is a family of dicots, dicotyledonous flowering plants found mostly in the tropics (two-thirds of the genera are from the New World tropics) comprising c. 175 genera and c. 5115 known species. Melastomes are annual or perennial h ...
(e.g. ''
Miconia ''Miconia'' is a genus of flowering plants in the glory bush family, Melastomataceae, native to warm temperate to tropical regions of the Americas. The species are mostly shrubs and small to medium-sized trees up to 15 m tall. The generic name ...
'' and ''
Tibouchina ''Tibouchina'' "Tibouchina." Webster's Third New International Dictionary of the English Language, Unabridged. Merriam Webster, 1961. is a Neotropical realm, neotropical flowering plant genus in the family Melastomataceae. Species of this genus a ...
'') and trees are generally overgrown with
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s and
hemiepiphyte A hemiepiphyte is a plant that spends part of its life cycle as an epiphyte. The seeds of primary hemiepiphytes germinate in the canopy and initially live epiphytically. They send roots downward, and these roots eventually make contact with t ...
s like ''
Coussapoa ''Coussapoa'' is a genus of flowering plants belonging to the family Urticaceae. Its native range is Mexico, Central & Southern Tropical America. Species: *''Coussapoa angustifolia'' *''Coussapoa arachnoidea'' *''Coussapoa argentea'' *''Co ...
'' (
Urticaceae The Urticaceae are a family, the nettle family, of flowering plants. The family name comes from the genus ''Urtica''. The Urticaceae include a number of well-known and useful plants, including nettles in the genus ''Urtica'', ramie (''Boehmeri ...
).


Behavior


Movement

The barred forest falcon is a year-round resident throughout its range but
vagrants Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
have been recorded beyond it.


Feeding

The barred forest falcon feeds on
lizards Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia althou ...
, small birds,
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s (mainly
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s and
marsupials Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a po ...
such as the
Brazilian slender opossum The Brazilian slender opossum (''Marmosops paulensis'') is an opossum species from South America. It is found in moist montane forest in the Atlantic Forest region of southeastern Brazil, including the states of Minas Gerais, Rio de Janeiro, S ...
, ''Marmosops paulensis''), bats, snakes, and frogs.Olmos, Fábio; Pacheco, José Fernando & Silveira, Luís Fábio (2006): Notas sobre aves de rapina (Cathartidae, Acciptridae e Falconidae) brasileiras
otes on Brazilian birds of prey High Laver is a village and civil parish in the Epping Forest district of the county of Essex, England. The parish is noted for its association with the philosopher John Locke. History High Laver is historically a rural agricultural parish, pred ...
''Revista Brasileira de Ornitologia'' 14(4): 401-404 ortuguese with English abstractbr>PDF fulltext
/ref> Like ''
Accipiter ''Accipiter'' is a genus of Bird of prey, birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (ex ...
''
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s, they often hunt prey by sitting quietly on tree branches and waiting for their victims to appear. When the prey arrives, the forest falcons quickly ambush them, attempting to catch them with a brief, flying pursuit. However, forest falcons also use other techniques to hunt prey, such as chasing prey on foot, following
army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limit ...
swarms, flushing prey from cover, and nest robbing. The species has also been recorded to snatch animals from traps or cages, for example during mark-recapture studies.


Breeding

The barred forest falcon's breeding season in Guatemala is from mid-April to early August. The season is not well known elsewhere but includes early June in Amazonian Brazil and October in Colombia. Almost all of the data on its breeding biology were collected in Guatemala during the Peregrine Fund's Maya Project. The species nests in tree cavities, almost always natural ones but sometimes those excavated by woodpeckers. Estimated heights were above the ground. Both nest territory and mate fidelity are high. The clutch size is two to six eggs and the female alone incubates. The incubation period is 33 to 35 days and time to fledging averages 38 days. Both parents provision the nestlings but the male provides about 2/3 of the effort.


Vocalization

The barred forest falcon's "advertising call" is "a somewhat dog-like, monotonous yapping, “keyak”, “ark” or similar". Another call is a "bouncing-ball series...“cah cah cah-cah-cah”, or similar". Neighboring males spar when one gives "a rapid, swelling chitter and the other a rapid clucking".


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the barred forest falcon as being of Least Concern. It has an extremely large range and an estimated population of at least a half million mature individuals, though the latter is believed to be decreasing. No immediate threats have been identified.


References


Further reading

* {{Taxonbar, from=Q1089154 barred forest falcon barred forest falcon Birds of the Amazon Basin Birds of Bolivia Birds of Brazil Birds of Colombia Birds of Central America Birds of Ecuador Birds of the Guianas Birds of Paraguay Birds of Venezuela barred forest falcon Taxa named by Louis Jean Pierre Vieillot Taxonomy articles created by Polbot