Banksia Xylothemelia
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''Banksia xylothemelia'' is a sprawling woody
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
of the family
Proteaceae The Proteaceae form a family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genera with about 1,660 known species. Together with the Platanaceae and Nelumbonaceae, they make up the order Pro ...
endemic to southern
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, one of the many species commonly known as dryandras and until recently called ''Dryandra xylothemelia''. To date it is almost unknown in cultivation.


Description

It grows as a sprawling shrub up to high, often with basal shoots arising from an underground lignotuber. It has
woolly Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. As an ...
stems. The leaves are
pinnatifid Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common Anatomical terms of location#Axes, axis. Pinnation occurs in biological morphology (biology), morphology, in Crysta ...
with five to nine leaflets on each side, long, wide, smooth above but woolly beneath. Flowers occur in dome shaped heads up to across, attached directly to an older stem, or on a short stalk. Flower heads contain from 80 to 100 bright yellow flowers. Each flower consists of a tubular
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when ...
made up of four fused
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, and one long wiry style. The styles are hooked rather than straight, and are initially trapped inside the upper perianth parts, but break free at
anthesis Anthesis is the period during which a flower is fully open and functional. It may also refer to the onset of that period. The onset of anthesis is spectacular in some species. In ''Banksia'' species, for example, anthesis involves the extension ...
. Seeds are produced in follicles embedded in the woody bases of the flower heads.


Taxonomy

Early collections of ''B. xylothemelia'' include a specimen collected in the Dragon Rocks Nature Reserve by Robert M. Buehrig on 9 December 1993, and a specimen collected from west of
Lake King Lake King is a town in the eastern Wheatbelt region of Western Australia, from Perth along State Route 40 between Kelmscott and Ravensthorpe. As of 2016, the town had a population of 95. The 2011 census recorded both the population of the tow ...
by P. G. Wilson. On 11 October 1994,
Alex George Alexander or Alex George may refer to: *Alex George (botanist) (born 1939), Australian botanist * Alexander L. George (1920–2006), American political scientist * Alexander George (philosopher), American philosopher *Alex George (motorcyclist), Sc ...
collected what would later become the
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
, from a location on the Holt Rock South Road, north of Newdigate-Lake King Road, at . Two years later, George published a formal description of the species in ''
Nuytsia ''Nuytsia floribunda'' is a hemiparasitic tree found in Western Australia. The species is known locally as moodjar and, more recently, the Christmas tree or Western Australian Christmas tree. The display of intensely bright flowers during the ...
'' 10(3), placing it in genus ''
Dryandra ''Banksia'' ser. ''Dryandra'' is a series of 94 species of shrub to small tree in the plant genus ''Banksia''. It was considered a separate genus named ''Dryandra'' until early 2007, when it was merged into ''Banksia'' on the basis of extensiv ...
'', subgenus ''Dryandra'' and series ''Armatae''. He gave it the specific name "xylothemelia", from the Greek "xylon" (''wood'') and "themelios" (''foundation''), in reference to species' lignotuber, the possession of which distinguishes it from its close relative '' B. cirsioides''; thus the full name of the species was for a time ''Dryandra xylothemelia'' A.S.George. Early in 2007, Austin Mast and Kevin Thiele transferred all ''Dryandra'' taxa to ''Banksia''. The current name for this species is therefore ''Banksia xylothemelia'' (A.S.George) A.R.Mast & K.R.Thiele. As an interim measure, Mast and Thiele placed all but one ''Dryandra'' taxon in ''Banksia'' ser. ''Dryandra''.


Distribution and habitat

It occurs in sand over
laterite Laterite is both a soil and a rock type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by ...
or in gravelly loam, amongst heath or sometimes
mallee woodlands and shrublands Mallee Woodlands and Shrublands is one of 32 Major Vegetation Groups defined by the Australian Government Department of the Environment and Energy. Description " Mallee" refers to the growth habit of a group of (mainly) eucalypt species that g ...
. It occurs only within the
Esperance Plains Esperance Plains, also known as Eyre Botanical District, is a biogeographic region in southern Western Australia on the south coast between the Avon Wheatbelt and Hampton bioregions, and bordered to the north by the Mallee region. It is a pl ...
and Mallee biogeographic region, between
Kulin Kulin may refer to: Places *Kulin, Western Australia, a small town in Australia ** Shire of Kulin, a local government area *Kulin, Iran, a village near Tehran *Kulin, Lower Silesian Voivodeship, a village in south-west Poland *Kulin, Kuyavian-Pome ...
, Lake Magenta,
Frank Hann National Park Frank Hann National Park is a national park in Western Australia, located east-southeast of the capital, Perth in the Shire of Lake Grace. It was named for Frank Hann, an early explorer of the district. The park contains a wide array of flora, ...
and Hyden.


Ecology

Like most other
Proteaceae The Proteaceae form a family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genera with about 1,660 known species. Together with the Platanaceae and Nelumbonaceae, they make up the order Pro ...
, ''B. xylothemelia'' has proteoid roots, roots with dense clusters of short lateral rootlets that form a mat in the soil just below the leaf litter. These enhance solubilisation of nutrients, thus allowing nutrient uptake in low-nutrient soils such as the
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
-deficient native soils of Australia. It has a lignotuber, so is able to resprout after being burned to the ground by bushfire. It has been given a rating of "Priority Three – Poorly Known Taxa" on Western Australia's Department of Environment and Conservation's Declared Rare and Priority Flora List. An assessment of the potential
impact of climate change The effects of climate change impact the physical environment, ecosystems and human societies. The environmental effects of climate change are broad and far-reaching. They affect the water cycle, oceans, sea and land ice (glaciers), sea level ...
on this species found that its range is likely to contract by half under mild change, and severe change is likely to lead to
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
unless it migrates rapidly into newly habitable areas.


Cultivation

''Banksia xylothemelia'' is rarely cultivated. It prefers
lateritic Laterite is both a soil and a rock type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by ...
and other heavy soils, and tolerates dry conditions and moderate frosts once established. Tony Cavanagh and
Margaret Pieroni Margaret Pieroni is a Western Australian botanical artist Botanical illustration is the art of depicting the form, color, and details of plant species, frequently in watercolor paintings. They must be scientifically accurate but often also ha ...
suggest that it would be best suited to a small garden in a warm sunny area. Seed takes six to seven weeks to germinate; germination rates appear to be quite low, with most reports suggesting success rates of less than 30%.


References


External links

* * * {{Taxonbar, from=Q4856766 xylothemelia Endemic flora of Western Australia Taxa named by Kevin Thiele