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''Banksia nivea'', commonly known as honeypot dryandra, is a species of rounded shrub that is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to Western Australia. The
Noongar The Noongar (, also spelt Noongah, Nyungar , Nyoongar, Nyoongah, Nyungah, Nyugah, and Yunga ) are Aboriginal Australian peoples who live in the south-west corner of Western Australia, from Geraldton on the west coast to Esperance on the so ...
peoples know the plant as bulgalla. It has linear, pinnatipartite leaves with triangular lobes, heads of cream-coloured and orange or red flowers and
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
, egg-shaped follicles.


Description

''Banksia nivea'' is a rounded, much-branched shrub that typically grows to high and wide but does not form a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
. It has linear, pinnatipartite leaves that are long and wide on a petiole long. There are between 45 and 85 triangular lobes on each side of the leaves. Between seventy and ninety cream-coloured and orange or red flowers are borne in head on the ends of branches with oblong to egg-shaped involucral bracts long at the base of the head. The
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when ...
is long and the
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
long. Flowering occurs in April or from July to November and the follicles are egg-shaped, long and almost glabrous.


Taxonomy and naming

''Banksia nivea'' was first collected by
Jacques Labillardière Jacques-Julien Houtou de Labillardière (28 October 1755 – 8 January 1834) was a French biologist noted for his descriptions of the flora of Australia. Labillardière was a member of a voyage in search of the Jean-François de Galaup, comte ...
in the vicinity of
Esperance Bay Esperance Bay is a bay on the south coast of Western Australia. Nominally located at , it is the site of the town of Esperance. The bay was discovered on 9 December 1792 by a French expedition under Bruni d'Entrecasteaux, which sailed in searc ...
between 15 and 17 December 1792, during a search for the naturalist
Claude Riche Claude-Antoine-Gaspard Riche (20 August 1762 – 5 September 1797) was a naturalist on Bruni d'Entrecasteaux's 1791 expedition in search of the lost ships of Jean-François de Galaup, comte de La Pérouse. Cape Riche, on the south coast of Austra ...
, who had become lost on the Australian mainland. Labillardière formally described and figured the species in ''
Relation du Voyage à la Recherche de la Pérouse ''Relation du Voyage à la Recherche de la Pérouse'' is an 1800 book that gives an account of the 1791-1793 d'Entrecasteaux expedition to Australasia. The title refers to the search for La Pérouse, who disappeared in the region in 1788, a popu ...
,'' his account of the voyage published in 1800. In 1810 Robert Brown transferred it into a new genus, ''
Dryandra ''Banksia'' ser. ''Dryandra'' is a series of 94 species of shrub to small tree in the plant genus ''Banksia''. It was considered a separate genus named ''Dryandra'' until early 2007, when it was merged into ''Banksia'' on the basis of extensiv ...
'' as ''D. nivea''. In 1996, Alex George described two subspecies of ''Dryandra nivea'': * ''Dryandra nivea'' (Labill.) R.Br. var. ''nivea'' that has a pistil long and leaves wide; * ''Dryandra nivea'' var. ''uliginosa'' A.S.George that has a pistil long and leaves wide. In 2007
Austin Mast Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently a professor within the Department of Biological Science at Florida State University (FSU), and has been dir ...
and
Kevin Thiele Kevin R. Thiele is currently an adjunct associate professor at the University of Western Australia and the director of Taxonomy Australia. He was the curator of the Western Australian Herbarium from 2006 to 2015. His research interests include ...
transferred all ''Dryandra'' species into ''Banksia'', reinstating Labillardière's ''Banksia nivea'' and renaming the two subspecies ''B. nivea'' Labill. subsp. ''nivea'' and ''B. nivea'' (A.S.George) A.R.Mast & K.R.Thiele subsp. ''uliginosa'', the names accepted by the
Australian Plant Census The Australian Plant Census (APC) provides an online interface to currently accepted, published, scientific names of the vascular flora of Australia, as one of the output interfaces of the national government Integrated Biodiversity Information Syst ...
. A third subspecies (''B. nivea'' subsp. ''Morangup (M.Pieroni 9/42'') WA Herbarium) has been named but not yet formally described.


Distribution and habitat

Honeypot dryandra is widespread between Lake Indoon (near Eneabba), Ongerup and
Israelite Bay Israelite Bay is a bay and locality on the south coast of Western Australia. Situated in the Shire of Esperance local government area, it lies east of Esperance and the Cape Arid National Park, within the Nuytsland Nature Reserve and the Grea ...
. Subspecies ''nivea'' grows in woodland and
kwongan Kwongan is plant community found in south-western Western Australia. The name is a Bibbelmun (Noongar) Aboriginal term of wide geographical use defined by Beard (1976) as Kwongan has replaced other terms applied by European botanists such as ...
Subspecies ''uliginosa'' has a narrow distribution from east of Busselton and on the
Scott River The Scott River is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map accessed March 9, 2011 river in Siskiyou County, California, United States. It is a tributary of the Klamath River, one of the ...
plain where it grows in thick scrub.


Ecology

Species of
nectarivorous In zoology, a nectarivore is an animal which derives its energy and nutrient requirements from a diet consisting mainly or exclusively of the sugar-rich nectar produced by flowering plants. Nectar as a food source presents a number of benefits a ...
birds that have been observed feeding on ''B. nivea'' include '' Acanthorhynchus superciliosus'' (western spinebill). Black cockatoos have also been recorded feeding upon the seed, though it is not clear which species of black cockatoo was observed, '' Calyptorhynchus baudinii'' (Baudin's black cockatoo) or '' C. latirostris'' (Carnaby's black cockatoo).


Conservation status

Subspecies ''nivea'' is classified as "not threatened" by the Western Australian Government
Department of Parks and Wildlife The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the ''Conservation and Land Management Act 1984'' and implementing the state's conservation and e ...
, but subsp. ''uliginosa'' is classified as " Threatened Flora (Declared Rare Flora — Extant)" by the
Department of Environment and Conservation (Western Australia) The Department of Environment and Conservation (DEC) was a department of the Government of Western Australia that was responsible for implementing the state's conservation and environment legislation and regulations. It was formed on 1 July 2006 ...
.


References

* {{Taxonbar, from=Q27897796, from2=Q4856652
nivea Nivea (, stylized as NIVEA) is a German personal care brand that specializes in skin and body care. It is owned by the Hamburg-based company Beiersdorf Global AG. The company was founded on 28 March 1882, by Paul Carl Beiersdorf. In 1890, it w ...
Endemic flora of Western Australia Eudicots of Western Australia Taxa named by Jacques Labillardière Plants described in 1800