Balea Sarsii
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''Balea sarsii'' is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of air-breathing
land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial gastropod mollusks that have shells (those without shells are known as ...
, a
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate Pulmonata or pulmonates, is an informal group (previously an order, and before that a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group includ ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
in the family
Clausiliidae Clausiliidae, also known by their common name the door snails, are a taxonomic family of small, very elongate, mostly left-handed, air-breathing land snails, sinistral terrestrial pulmonate gastropod mollusks.MolluscaBase eds. (2020). MolluscaBa ...
. The species has long been overlooked because of confusion with ''
Balea perversa ''Balea perversa'', also known as the wall snail or tree snail, is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Clausiliidae, the door snails. The shell of this species is left-handed in coiling ...
''.


Taxonomy

The specific name ''sarsii'' is in honour of Norwegian biologist
Michael Sars Michael Sars (30 August 1805 – 22 October 1869) was a Norwegian theologian and biologist. Biography Sars was born in Bergen, Norway. He studied natural history and theology at Royal Frederick University from 1823 and completed a cand.theol. de ...
. According to Welter-Schultes ''Balea heydeni'' is conspecific with ''Balea sarsii''. The name ''heydenii'' has now been superseded as the species was described from Norway at an earlier date than von Maltzan, under the name ''sarsii'' Pfeiffer (synonym ''lucifuga'' Bourguignat 1857). This species has been separated from the closely related, common and wide-spread ''
Balea perversa ''Balea perversa'', also known as the wall snail or tree snail, is a species of air-breathing land snail, a terrestrial pulmonate gastropod mollusk in the family Clausiliidae, the door snails. The shell of this species is left-handed in coiling ...
'' and redescribed by Gittenberger et al. (2006)Gittenberger E., Prece R. C. & Ripken T. E. J. (2006). "''Balea heydeni'' von Maltzan, 1881 (Pulmonata: Clausiliidae): an overlooked but widely distributed European species". ''
Journal of Conchology The ''Journal of Conchology'' is a peer-reviewed scientific journal published by The Conchological Society of Great Britain and Ireland, covering research in conchology and malacology. It claims to be the world's oldest continuing publication on t ...
'' 39(2): 145-150.
under its junior synonym name ''Balea heydeni'' von Maltzan, 1881. The valid name for this taxon is, however, ''Balea sarsii''.


Distribution

''Balea sarsii'' is a western-Atlantic element, found on the Atlantic Islands, in Portugal, northwestern Spain, and the coastal parts of France, Belgium and the Netherlands. In Britain and Ireland it is the commoner of the two ''Balea'' species, occurring also in the inland. The only locality in Denmark is Møns Klint. A recently undertaken revision of ''Balea''-material from Sweden and Norway in the Göteborg Natural History Museum and the
Swedish Museum of Natural History The Swedish Museum of Natural History ( sv, Naturhistoriska riksmuseet, literally, the National Museum of Natural History), in Stockholm, is one of two major museums of natural history in Sweden, the other one being located in Gothenburg. The ...
in Stockholm revealed two localities for ''Balea sarsii'' from the Swedish west coast (the Island of
Vinga Vinga is a commune in Arad County, western Romania, south of the county seat of Arad, with a population of 5,828 inhabitants (as of 2011). Vinga is located in the northern section of the Banat. The people in Vinga are mainly Romanians, the sec ...
outside Göteborg and the Island Storön in the archipelago of Väderöarna in the province of
Bohuslän Bohuslän (; da, Bohuslen; no, Båhuslen) is a Swedish province in Götaland, on the northernmost part of the country's west coast. It is bordered by Dalsland to the northeast, Västergötland to the southeast, the Skagerrak arm of the North Sea ...
). Totally the species is now known from six Norwegian localities, of which five are situated in Hordaland County. It is considered a very rare species in Norway, because only sixteen Norwegian specimens have been found, among thousands of ''Balea perversa''. This species is known to occur in: * Atlantic Islands *
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
* northwestern
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
* coastal parts of
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
* coastal parts of
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
* coastal parts of
the Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
*
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...
*
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
*
Møns Klint Møns Klint is a 6 km stretch of limestone and chalk cliffs along the eastern coast of the Danish island of Møn in the Baltic Sea. Some of the cliffs fall a sheer 120 m to the sea below. The highest cliff is , which is 128 m above sea lev ...
in
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...
*
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
*
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and t ...
:
Hordaland Hordaland () was a county in Norway, bordering Sogn og Fjordane, Buskerud, Telemark, and Rogaland counties. Hordaland was the third largest county, after Akershus and Oslo, by population. The county government was the Hordaland County Municipal ...
County and
Sogn og Fjordane Sogn og Fjordane (; English: "Sogn and Fjordane") was, up to 1 January 2020, a county in western Norway, when it was merged to become part of Vestland county. Bordering previous counties Møre og Romsdal, Oppland, Buskerud, and Hordaland, the cou ...
County The type locality is
Florø is a town and the administrative centre of the municipality of Kinn, Vestland, Norway. The town was founded by royal decree in 1860 as a ladested on the island of Florelandet, located between the Botnafjorden and Solheimsfjorden. Florø is No ...
in
Sogn og Fjordane Sogn og Fjordane (; English: "Sogn and Fjordane") was, up to 1 January 2020, a county in western Norway, when it was merged to become part of Vestland county. Bordering previous counties Møre og Romsdal, Oppland, Buskerud, and Hordaland, the cou ...
County. This is also the northernmost locality of ''Balea sarsii'' known in Europe.


Description

The shell of this species is sinistral (left-handed) in its coiling. Differs from ''Balea perversa'' in its more slender and yellowish rather than brownish shell. The first whorl increases in diameter more rapidly and the sculpture is more weakly striated (with coarse growth lines rather than distinct riblets). ''Balea sarsii'', differs from ''Balea perversa'' through its, in most cases, shorter, broader shell, in which the whorls increase more quickly in width. ''Balea sarsii'' reaches a maximal height of 7 mm; ''Balea perversa'' might be higher (up to 10 mm). The
sculpture Sculpture is the branch of the visual arts that operates in three dimensions. Sculpture is the three-dimensional art work which is physically presented in the dimensions of height, width and depth. It is one of the plastic arts. Durable sc ...
in ''Balea sarsii'' is also more wrinkled, coarser and less regular, compared to the fine, regular riblets typical for ''Balea perversa''. The shell colour is also yellowish – light brown, compared to the usually somewhat darker brown in ''Balea perversa''. A weak, parietal denticle may be present in ''Balea perversa'', but not in ''Balea sarsii''. All these characters are, however, variable; especially the shell dimensions. The most reliable character to use in separating the two species is the form of the
apical Apical means "pertaining to an apex". It may refer to: *Apical ancestor, refers to the last common ancestor of an entire group, such as a species (biology) or a clan (anthropology) *Apical (anatomy), an anatomical term of location for features loc ...
whorls: in ''Balea perversa'' this part of the shell is almost cylindrical, but in ''Balea sarsii'' it is conical.


Ecology

''Balea sarsii'' sometimes co-occur with ''Balea perversa''. Such
sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...
occurrences are not uncommon in various parts of the distribution area, and probably both species have very similar ecology. ''Balea''-species are rarely found on the ground, they are climbers and live on vertical surfaces (rocks, trees etc.)


References

This article incorporates CC-BY-3.0 text from the reference


External links


''Balea sarsii''
at
Animalbase AnimalBase is a project brought to life in 2004 and is maintained by the University of Göttingen, Germany. The goal of the AnimalBase project is to digitize early zoological literature, provide copyright-free open access to zoological works, and pr ...
{{Taxonbar, from=Q4850669 Clausiliidae Molluscs of Europe Gastropods described in 1847 Taxobox binomials not recognized by IUCN