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:''Common names: Trans-Pecos copperhead, western copperhead'' ''Agkistrodon contortrix pictigaster'' was formerly a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper subspecies found in the
Trans-Pecos The Trans-Pecos, as originally defined in 1887 by the Texas geologist Robert T. Hill, is the portion of Texas that lies west of the Pecos River. The term is considered synonymous with Far West Texas, a subdivision of West Texas. The Trans-Peco ...
region of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
in western
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
, and northeastern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. However, recent taxonomic changes do not recognize the Trans-Pecos copperhead (''A. c. pictigaster'') as a valid taxon.


New Taxonomy

The Trans-Pecos copperhead (''Agkistrodon contortrix pictigaster'') Conant R. 1975. ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America''. Second Edition. First published in 1958. Houghton Mifflin Company Boston. 429 pp. 48 plates. (hc), (pb). was once classified as a subspecies of the copperhead (''
Agkistrodon contortrix The eastern copperhead (''Agkistrodon contortrix''), also known as the copperhead, is a species of venomous snake, a pit viper, endemic to eastern North America; it is a member of the subfamily Crotalinae in the family Viperidae. The eastern co ...
''). However, DNA based studies published in 2008 and 2015, revealed no significant genetic difference between the Trans-Pecos copperhead (''Agkistrodon contortrix pictigaster'') and the broad-banded copperhead, (''Agkistrodon contortrix laticinctus''). The two subspecies were synonymized and elevated to full species, with the oldest published name, ''
Agkistrodon laticinctus ''Agkistrodon laticinctus'', commonly known as the broad-banded copperhead, is a venomous pit viper species,Frank T. Burbrink, Timothy J. Guiher (2014). Considering gene flow when using coalescent methods to delimit lineages of North American pi ...
'', having priority. The resulting taxonomy does not recognize the Trans-Pecos copperhead (''Agkistrodon contortrix pictigaster'') as a valid taxon.Guiher TJ, Burbrink FT (2008). ''Demographic and phylogeographic histories of two venomous North American snakes of the genus Agkistrodon.'' Molecular Phylogenetics and Evolution, 48: 543–553.Burbrink, Frank T. and Timothy J. Guiher. 2014. ''Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon.'
Zoological Journal of the Linnean Society, 173: 505–526
Several subsequent reviews and species accounts have followed and supported the revised taxonomy.Crother, B. I. ( editor). 2017. ''Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding, 8th. edition.'' SSAR Herpetological Circular 43, 1–102 pp. (page 59) Powell, Robert, Roger Conant, and Joseph T. Collins. 2016. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, 4th ed. Houghton Mifflin Harcourt Publishing Co. New York. 494 pp. Uetz P, Freed P, Aguilar R, Hošek J (editors) (2021). The Reptile Database,
Agkistrodon laticinctus
' (accessed 30 August 2021)
Information on this snake can be found in the ''
Agkistrodon laticinctus ''Agkistrodon laticinctus'', commonly known as the broad-banded copperhead, is a venomous pit viper species,Frank T. Burbrink, Timothy J. Guiher (2014). Considering gene flow when using coalescent methods to delimit lineages of North American pi ...
'' article.


Description

This subspecies is typically a light tan in color, with darker brown, wide crossbands. Their actual color varies by locality, varying from a red-brown, to a gray-brown. The species can be difficult to distinguish from the broad-banded copperhead, '' A. c. laticinctus''. The only notable physical difference between the subspecies is that the Trans-Pecos copperhead tends to have an elaborately patterned underside, often being an irregular, white and black pattern whereas the broad-banded tends to be plain white, only have minimal patterning, or have elongated random blotching instead of a distinct pattern. The subspecies
intergrade In zoology, intergradation is the way in which two distinct subspecies are connected via areas where populations are found that have the characteristics of both. There are two types of intergradation: primary and secondary intergradation. Primary ...
where their ranges overlap, further confusing identification. It is easily distinguished from other subspecies of copperhead, in that other species typically have banding that narrows at the spine, creating hourglass shapes, whereas ''A. c. pictigaster'' has bands that do not narrow at the spine. They grow to approximately 20-36 inches (50–90 cm) in length. As juveniles, all species of ''
Agkistrodon ''Agkistrodon'' is a genus of venomous pit vipers commonly known as American moccasins.Crother, B. I. (ed.). 2017. ''Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confi ...
'' have a bright green-yellow color to their
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammal ...
tip believed to be used as a lure to attract prey items to approach within striking range. The color fades to a grey or brown at about a year of age.


Behavior

Like all ''A. contortrix'' subspecies, ''A. c. pictigaster'' is a nocturnal ambush
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
. Their
diet Diet may refer to: Food * Diet (nutrition), the sum of the food consumed by an organism or group * Dieting, the deliberate selection of food to control body weight or nutrient intake ** Diet food, foods that aid in creating a diet for weight loss ...
consists of primarily lizards,
frog A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is ...
s, and
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s. Their choice of habitat is the driest among the copperhead subspecies, preferring rocky, lightly vegetated, canyon areas. They are
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, giving birth to a litter of up to 8 young in the early fall.


Venom

Like most pit viper species, ''A. contortrix'' has a
hemotoxic Hemotoxins, haemotoxins or hematotoxins are toxins that destroy red blood cells, disrupt blood clotting, and/or cause organ degeneration and generalized tissue damage. The term ''hemotoxin'' is to some degree a misnomer since toxins that damage t ...
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a st ...
, which is delivered through hinged, hollow fangs set in the front of their jaw. The fangs work like
hypodermic needle A hypodermic needle (from Greek ὑπο- (''hypo-'' = under), and δέρμα (''derma'' = skin)), one of a category of medical tools which enter the skin, called sharps, is a very thin, hollow tube with one sharp tip. It is commonly used w ...
s, injecting the venom in a single, quick striking motion. Copperhead venom is not considered to be life-threatening to an otherwise healthy adult, but it can cause localized swelling, necrosis, and severe pain. Any bite from a venomous snake should be considered serious and medical treatment sought. There is no
antivenin Antivenom, also known as antivenin, venom antiserum, and antivenom immunoglobulin, is a specific treatment for envenomation. It is composed of antibodies and used to treat certain venomous bites and stings. Antivenoms are recommended only if th ...
specifically manufactured for copperheads, but CroFab, which makes use of the venom from the cottonmouth ('' Agkistrodon piscivorus''), can be used in cases of a severe envenomation.


References


External links

*
''Agkistrodon contortrix''
a
University of Texas
Accessed 7 December 2007.

a
WhoZoo
Accessed 7 December 2007. {{Taxonbar, from=Q11684612 contortrix pictigaster Reptiles of the United States