Australonycteris
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''Australonycteris'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
and
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus of microchiropteran
bat Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most bi ...
with the single species ''Australonycteris clarkae''. The species is known from fragmentary remains found at the
Murgon fossil site The Murgon fossil site is a paleontology, paleontological site of early Eocene age in south-eastern Queensland, Australia. It lies near the town of Murgon, some 270 km north-west of Brisbane. The Murgon site is important as the only site on ...
, in south-eastern
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
, dating to the early
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
, 54.6 million years ago. It is the oldest bat from the Southern Hemisphere and one of the oldest bats in the world, and inhabited forests and swampy areas, with a diet of insects and possibly small fish.


Taxonomy

A monotypic genus allied to the family
Archaeonycteridae Archaeonycteridae (formerly spelled Archaeonycterididae) is a Family (biology), family of extinct bats. It was originally erected by the Swiss people, Swiss naturalist Pierre Revilliod as Archaeonycterididae to hold the genus ''Archaeonycteris''. ...
, or classified by an indeterminate familial arrangement, describing
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
material collected at Murgon in 1994. The type specimen is a tooth. The material was discovered at the Tingamarra Local Fauna – Boat Mountain deposit, the type and only known location of fossil evidence of the species. The phylogenetic relationship to other Chiroptera is uncertain, but may represent an early geographic dispersal and separation from the crown clade of bat taxa found on other continents. The genus name is derived from ancient Greek, indicating a bat with an Australian distribution. The specific epithet honours Elaine Clark, a person closely associated with the palaeontological research at Riversleigh and Murgon.


Description

''Australonycteris clarkae'', one of the earliest bats in the fossil record, is known from several upper and lower teeth, an edentulous lower-jaw fragment, a partial periotic bone, and several postcranial fragments. It has a forearm length of , making it a medium-size bat species, and it could echolocate. ''Australonycteris'' displayed some differences in dental anatomy, compared to extant bats. Unlike other purported Eocene bats, such as the '' Wyonycteris'' species of Wyoming and Europe in the northern hemisphere, ''Australonycteris clarkae'' possessed many of the anatomical characteristics of the modern microchiropteran. The tooth tip of ''A. clarkae'' shows wear consistent with carrying prey with a hard carapace, such as beetles, or bony skeleton of fish; the body size of the species would make predation of fish at least possible. ''Australonycteris'' is thought to have been fully capable of flight.


See also

* Icaronycteris


References

* * {{Taxonbar, from=Q2173043 Eocene mammals of Australia Eocene bats Prehistoric bat genera Prehistoric monotypic mammal genera Fossil taxa described in 1994