Australoheros Facetus
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''Australoheros facetus'', the chameleon cichlid or chanchito, is a species of
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Cichlasomatinae The Cichlasomatinae are a subfamily of cichlid fishes, including all ciclids native to the Greater Antilles (Cuba and Hispaniola), United States (southern Texas), Mexico and Central America, and many of the cichlids from South America (the other ...
which is native to northern Argentina, Paraguay, Uruguay and southern Brazil.


Description

''Australoheros facetus'' has a high and deeply compressed body which is covered with large scales. It has a small, upward pointing mouth in which the jaws do not reach farther than the forward margin of the eye. It has a long
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
which extends to two thirds of the total length of the fish. The
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
has a rounded shape. It is distinguished from other species in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
'' Australoheros'' by its longer lower jaw, upward pointing mouth and in having the shortest dorsal scale cover and the fewest scales on the dorsal and
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
s. Most specimens, 80% of fish, have four abdominal bars while these are present in only half of the individuals of its congeners. They normally attain in length but males have been recorded at The colour varies from brassy yellow through to greenish and even black and it is normally marked with a number of dark transverse bands. The males are larger than the females with larger fins.


Distribution

''Australoheros facetus'' has a native range in the Río de la Plata Basin in Argentina, Brazil, Paraguay and Uruguay, and in the coastal drainages of Uruguay,
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
in Brazil and
Buenos Aires Buenos Aires ( or ; ), officially the Autonomous City of Buenos Aires ( es, link=no, Ciudad Autónoma de Buenos Aires), is the capital and primate city of Argentina. The city is located on the western shore of the Río de la Plata, on South ...
in Argentina. It has been introduced and established to southern
Iberia The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
and Chile. It is also known to have been introduced to Germany and the Philippines but it failed to establish in Germany and its status in the Philippines is not known.


Habitat and biology

''Australoheros facetus'' is a freshwater species which occurs in creeks, rivers, swamps and lakes. It is a diurnal species which is omnivorous and rather opportunistic in its feeding habits. Its diet is known to include
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...
and plants and it preys on small aquatic animals, even small fish, but mainly
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s and insect larvae. It demonstrates an ability to tolerate low temperatures. ''A. facetus'' has been recorded as a host of the
trematode Trematoda is a class of flatworms known as flukes. They are obligate internal parasites with a complex life cycle requiring at least two hosts. The intermediate host, in which asexual reproduction occurs, is usually a snail. The definitive host ...
''
Centrocestus formosanus ''Centrocestus formosanus'' is a trematode parasite of Asian origin that has found its way into North American streams and rivers. It not only affects the fountain darter but many species of commercially important fishes. It is also capable of i ...
'', an intestinal
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
of
piscivorous A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...
birds and mammals, which is known to infect people in its native range which is Asia. Chameleon cichlids lay their eggs on a stone or piece of wood in the open and the eggs and fry are cared for by both parents for up to three weeks, with the eggs hatching after 2–4 days. In the rivers it has been introduced to in Europe it breeds spawns when the temperature climbs to more than . Pairs are formed for spawning and both sexes are
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
. In Iberia breeding occurs in the months of April to June.


Name

The
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
was collected by
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended fr ...
during his time on board
H.M.S. Beagle HMS ''Beagle'' was a 10-gun brig-sloop of the Royal Navy, one of more than 100 ships of this class. The vessel, constructed at a cost of £7,803 (roughly equivalent to £ in 2018), was launched on 11 May 1820 from the Woolwich Dockyard on th ...
and the species was described as ''Chromis facetus'' by Leonard Jenyns in 1842 in ''The zoology of the voyage of the H. M. S. Beagle, under the command of Captain Fitzroy, R. N., during the years 1832 to 1836''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "well made" or "fine" and Jenyns does not explain why he chose this name. The English
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
, chameleon cichlid, seems to originate in Brazil where this species is called , the "chameleon fish", as it was one of the first fishes to be observed to have an ability to change its colour. The alternative common name, ''chanchito'' is Spanish and means "piglet" and refers to its pugnacious nature and habit of digging up plants.


Aquaria

The chameleon cichlid is tolerant of a wide variety of water conditions and was one of the first species to be widespread in the hobby of fishkeeping and they are easy to breed.


References

{{Taxonbar, From=Q3576 Heroini Fish described in 1842