The Aulacidae are a small, cosmopolitan family of
wasp
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. ...
s, with two extant genera containing some 200 known species. They are primarily
endoparasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionary strategies within parasiti ...
s of wood wasps (
Xiphydriidae
Xiphydriidae are a family of wood wasps that includes around 150 species. They are located all over the world including North and South America, Australia, Europe, and others. Xiphydriidae larvae are wood borers in dead trees or branches of a ran ...
) and
xylophagous
Xylophagy is a term used in ecology to describe the habits of an herbivorous animal whose diet consists primarily (often solely) of wood. The word derives from Greek ''ξυλοφάγος'' (''xulophagos'') "eating wood", from ''ξύλον'' (') ...
beetle
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s (
Cerambycidae
The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
and
Buprestidae). They are closely related to the family
Gasteruptiidae, sharing the feature of having the first and second
metasoma
The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circul ...
l
tergites
A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'mar ...
fused, and having the head on a long
pronotal "neck", though they are not nearly as slender and elongate as gasteruptiids, nor are their hind legs club-like, and they have more sculptured
thoraces. They share the evanioid trait of having the metasoma attached very high above the hind
coxae on the
propodeum
The propodeum or propodium is the first abdominal segment in Apocrita Hymenoptera (wasps, bees and ants). It is fused with the thorax to form the mesosoma. It is a single large sclerite
A sclerite (Greek , ', meaning " hard") is a hardened bod ...
.
While generally rarely collected, they can be locally abundant in areas undergoing logging or forest fires. The rich fossil record of Aulacidae indicates they were quite abundant in the
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
.
References
Tree of Life AulacidaeCatalogus Evaniidorum
Further reading
*Jennings, J.T. & Austin, A.D. 2004, Biology and host relationships of aulacid and gasteruptiid wasps (Hymenoptera: Evanioidea): a review. In: Rajmohana, K., Sudheer, K., Girish Kumar, P., & Santhosh, S. (Eds) Perspectives on Biosystematics and Biodiversity. (University of Calicut, Kerala, India), pp. 187–215.
*Turrisi, G.F., Jennings, J.T. & Vilhelmsen, L. 2009, Phylogeny and generic concepts in the parasitoid wasp family Aulacidae (Hymenoptera: Evanioidea). Invertebrate Systematics 23, 27–59.
Evanioidea
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