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''Athertonia'' is a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of plants in the family
Proteaceae The Proteaceae form a family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genera with about 1,660 known species. Together with the Platanaceae and Nelumbonaceae, they make up the order Pro ...
. The sole described
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
is ''Athertonia diversifolia'', commonly known as Atherton oak, athertonia, creamy silky oak or white oak. It is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to a small part of the Wet Tropics of Queensland, Australia. A relative of the macadamia, it has potential in
horticulture Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
and the bushfood industry.


Description

''Athertonia diversifolia'' is a tree growing up to tall, the trunk may be fluted and may be buttressed. New shoots and young branches are densely covered in fine rust-coloured hairs. The leaf morphology is highly variable − from a simple elliptic shape to deeply
lobed The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
, and they may be with or without finely toothed margins. The overall leaf size also varies considerably, from up to . The petiole measures between long. The
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
is a pendulous
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
up to long, produced terminally, in the leaf
axils A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
, and on the branches. The fragrant flowers are cream to green with 4
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, and measure up to long. The fruit is green while developing − at maturity it is a blue (or occasionally rose coloured) lens-shaped
drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...
measuring around long, wide and thick. The hard, pitted endocarp contains a single seed.


Phenology

Flowering occurs from February to June, and fruit ripen around August to January.


Taxonomy

''Athertonia diversifolia'' was first described in 1918 as ''Helicia diversifolia'' by the Queensland Government botanist
Cyril Tenison White Cyril Tenison ("C.T.") White (17 August 1890 – 15 August 1950) was an Australian botanist. Early life White was born in Brisbane to Henry White, a trade broker, and Louisa ''nee'' Bailey. He attended school at South Brisbane State School, an ...
in his paper ''Contribution to the Queensland FLora'', published in the journal Botany Bulletin. Decades later, in 1955, the Dutch botanist
Hermann Otto Sleumer Hermann Otto Sleumer (February 21, 1906 in Saarbrücken – October 1, 1993 in Oegstgeest) was a Dutch botanist of German birth. The plant genera ''Sleumerodendron'' Virot (Proteaceae) and ''Sleumeria'' Utteridge, Nagam. & Teo (Icacinaceae The I ...
placed it in the genus ''
Hicksbeachia ''Hicksbeachia'' is a genus of two species of trees in the family Proteaceae. They are native to rainforests of northern New South Wales and southeastern Queensland. They are commonly known as red bopple nut or beef nut due to the bright red col ...
'', where it stayed until 1975 when Lawrie Johnson and
Barbara Briggs Barbara Gillian Briggs (born 1934) is one of the foremost Australian botanists. The '' IK'' lists 205 names of plants which have been published or co-published by her. She was one of the botanists in the Angiosperm Phylogeny Group, of th1998APG ...
revised the description and created a new genus, ''Athertonia'', for this plant.


Etymology

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
name is a reference to the
Atherton Tableland The Atherton Tableland is a fertile plateau which is part of the Great Dividing Range in Queensland, Australia. The principal river flowing across the plateau is the Barron River. It was dammed to form an irrigation reservoir named Lake Tina ...
s, where the
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
was collected. The species epithet is derived from the Latin words ''diverto'' (to differ), and ''folia'' (leaf), and refers to the varied leaf morphology it displays.


Vernacular names

The common names "Atherton oak", "silky oak" and "white oak" are given to this species as a reference to the similarity of its leaves to those of the unrelated English oak (''Quercus robur'').


Distribution and habitat

''Athertonia diversifolia'' is found in north Queensland from the
Cape Tribulation Cape Tribulation is a headland and coastal locality in the Shire of Douglas in northern Queensland, Australia. In the , Cape Tribulation had a population of 118 people. Geography The locality is north of Cairns. It is within the Daintree Na ...
area south to the
Atherton Tableland The Atherton Tableland is a fertile plateau which is part of the Great Dividing Range in Queensland, Australia. The principal river flowing across the plateau is the Barron River. It was dammed to form an irrigation reservoir named Lake Tina ...
s west of
Mount Bartle Frere Mount Bartle Frere (pronunciation mæɔnt̥ ˈbɐːɾəɫ ˈfɹɪə Ngajanji: Choorechillum) is the highest mountain in Queensland at an elevation of . The mountain was named after Sir Henry Bartle Frere, 1st Baronet, Sir Henry Bartle Frere ...
. It grows in rainforest, generally on deep soils of volcanic origin. The altitudinal range is mostly from around , but confirmed observations have been made in lowland forest at just above sea level in the
Cow Bay Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
area.


Ecology

The kernel contained within the hard endocarp of this species is edible to humans, and is also much sought after by native rats, which often gnaw through the hard shell and eat the kernel while it is still attached to the tree. The flowers attract a variety of nectar-eating birds.


Conservation

Much of the original habitat of the Atherton oak has been cleared, and what is left is highly fragmented. Despite this the species has been given the classification of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
, by both the
Queensland Government The Queensland Government is the democratic administrative authority of the Australian state of Queensland. The Government of Queensland, a parliamentary constitutional monarchy was formed in 1859 as prescribed in its Constitution, as amended fr ...
Department of Environment and Science (DES), and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN). Whilst the DES does not publish its procedure or qualification details, the IUCN's assessment states that the population of this species is "locally common and stable", and that the "area, extent and/or quality" of its habitat is not declining.


Uses

The Atherton oak has become widely cultivated, due to its attractive, glossy, lobed foliage, and its low maintenance.


Gallery

File:Athertonia-diversifolia-1-Australasian-Virtual-Herbarium-CC-BY.jpg, Ripe blue fruit File:Athertonia-diversifolia-4-Australasian-Virtual-Herbarium-CC-BY.jpg, Rose-coloured fruit File:Athertonia-diversifolia-2-Australasian-Virtual-Herbarium-CC-BY.jpg, Foliage File:Athertonia-diversifolia-5-Australasian-Virtual-Herbarium-CC-BY.jpg, The inflorescence File:Athertonia diversifolia 237508998.jpg, New growth File:Athertonia-diversifolia-SF23031-01.jpg, The endoncarp, after being gnawed by native rats to access the seed. File:Athertonia diversifolia 237508950.jpg, Underside of the leaf File:Athertonia diversifolia 224175498.jpg, Fruit with partially eaten kernel


References


External links

* *
View a map
of historical sightings of this species at the
Australasian Virtual Herbarium The ''Australasian Virtual Herbarium'' (AVH) is an online resource that allows access to plant specimen data held by various Australian and New Zealand herbaria. It is part of the Atlas of Living Australia (ALA), and was formed by the amalgamat ...

View observations
of this species on iNaturalist
View images
of this species on Flickriver {{Taxonbar, from1=Q5224696, from2=Q2869046 Proteaceae Trees of Australia Ornamental trees Endemic flora of Queensland Plants described in 1918 Proteales of Australia Monotypic Proteaceae genera Taxa named by Barbara G. Briggs Taxa named by Cyril Tenison White Wet Tropics of Queensland