''Atheris hispida'' is a

viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo, ...

. It is known for its extremely keeled

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...

that give it a bristly appearance. No

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are currently recognized. Common names include rough-scaled bush viper, spiny bush viper,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . hairy bush viper, and

others Others or The Others may refer to: Fictional characters * Others (A Song of Ice and Fire), Others (''A Song of Ice and Fire''), supernatural creatures in the fictional world of George R. R. Martin's fantasy series ''A Song of Ice and Fire'' * Ot ...

. Like all other vipers, it is

venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

Description

The males of this species grow to maximum total length of : body , tail . Females grow to a maximum total length of . The males are surprisingly long and slender compared to the females. The head has a short snout, more so in males than in females. The eyes are large and surrounded by 9–16

circumorbital scales In scaled reptiles, the ocular scales are those forming the margin of the eye.Wright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . The name originates from the term ''oculus'' which ...

. The orbits (eyes) are separated by 7–9 scales. The nostril is like a slit and separated from the eye by two scales. The eye and the

supralabials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...

are separated by a single row of scales. The supralabials number 7-10, of which the fourth is enlarged. The body is covered with elongated, heavily keeled dorsal scales that give this animal a unique 'shaggy' look, almost bristly in appearance. The scales around the head and neck are the longest, decreasing posteriorly. Midbody, the

are in 15–19 rows. There are 149–166

ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that conta ...

and 35–64

subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...

. The

anal scale Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving s ...

is single.

Common Names

Rough-scaled bush viper, spiny bush viper hairy bush viper, rough-scaled tree viper, African hairy bush viper, hairy viper.

Geographic range

Central and East Africa: northern and eastern

DR Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...

, southwestern

Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...

, west

Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

, and northwestern

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...

. The type locality given is "Lutunguru,

Kivu Kivu was the name for a large "region" in the Democratic Republic of the Congo under the rule of Mobutu Sese Seko that bordered Lake Kivu. It included three "Sub-Regions" ("Sous-Régions" in French): Nord-Kivu, Sud-Kivu and Maniema, correspondin ...

" (DR Congo). More specifically, Spawls & Branch (1995) describe the distribution as isolated populations in Kivu and Orientale Provinces in DR Congo, southeastern Ruwenzori in Uganda and the

Kakamega Forest Kakamega Forest is a tropical rainforest situated in the Kakamega County, Kakamega and Nandi_County, Nandi County of Kenya, northwest of the capital Nairobi, and near to the border with Uganda. It is Kenya's only tropical rainforest and is said to ...

in western Kenya.

Behavior

Capable of climbing reeds and stalks, this species is often found basking on top of flowers and terminal leaves. Mostly nocturnal.

Feeding

Feeds on

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...

s, and sometimes

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

s. Sometimes hunts for mammalian prey on the ground.

Reproduction

Females give birth to up to 12 young at a time. Newborns are about in total length.

Venom

Not much is known about their venom except that it is mainly

neurotoxic Neurotoxicity is a form of toxicity in which a biological, chemical, or physical agent produces an adverse effect on the structure or function of the central and/or peripheral nervous system. It occurs when exposure to a substance – specificall ...

. Besides the neurotoxins, they also carry

cytotoxins Cytotoxicity is the quality of being toxic to cells. Examples of toxic agents are an immune cell or some types of venom, e.g. from the puff adder (''Bitis arietans'') or brown recluse spider (''Loxosceles reclusa''). Cell physiology Treating cell ...

and fasciculins. Toxicity of individual specimens within the same species and subspecies can vary greatly based on several factors, including geographical region. Even the weather and altitude can influence toxicity (Ernst and Zug et al. 1996). A bite can be fatal to humans without access to proper

first aid First aid is the first and immediate assistance given to any person with either a minor or serious illness or injury, with care provided to preserve life, prevent the condition from worsening, or to promote recovery. It includes initial in ...

and subsequent

antivenom Antivenom, also known as antivenin, venom antiserum, and antivenom immunoglobulin, is a specific treatment for envenomation. It is composed of antibodies and used to treat certain venomous bites and stings. Antivenoms are recommended only if th ...

treatment. Until recently, their venom has often been regarded as less toxic than that of many other species, perhaps because bites are uncommon, but this turned out not to be the case. There are now a number of reports of bites that have led to severe hemorrhaging of internal organs.