''Astroconodon'' is an extinct genus of mammal from the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. Part of

Eutriconodonta Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the ...

, it was a small sized

predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...

, either a terrestrial

insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...

and

carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...

, or a semi-aquatic

piscivore A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...

. It is the first

eutriconodont found.

Description

The type species is ''A. denisoni''. Known from the

Antlers Formation The Antlers Formation is a stratum which ranges from Arkansas through southern Oklahoma into northeastern Texas. The stratum is thick consisting of silty to sandy mudstone and fine to coarse grained sandstone that is poorly to moderately sorted. ...

, its type specimen, ''FMNH PM 542'', was first described by Bryan Patterson in 1951. It is a generally rather common species, known from a large quantity of isolated teeth, exhibiting a high degree of variability. Its molars are on average 0.59 to 2

millimeter 330px, Different lengths as in respect to the electromagnetic spectrum, measured by the metre and its derived scales. The microwave is between 1 meter to 1 millimeter. The millimetre (American and British English spelling differences#-re, -er, ...

s long, and the

distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

ones see a marked increase in crown height. A second species, ''A. delicatus'', was later found in the

Cedar Mountain Formation The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid-Cretaceous. The formation was named for Cedar Mountain (Utah), Cedar Mountain in northern Emery Coun ...

. Its type locality is Mussentuchit (OMNH V239). It is smaller than ''A. denisoni'' by approximately 80%, and it differs from it, and most North American triconodontids, by lacking a lingual cingulid on the lower molars and premolars. A third species, currently unnamed, is known from the

Twin Mountains Formation The Twin Mountains Formation, also known as the Twin Mak Formation, is a sedimentary rock formation, within the Trinity Group, found in Texas of the United States of America. It is a terrestrial formation of Aptian age (Lower Cretaceous), and ...

. Not much has been said about it.

Phylogeny

Always identified as a "

triconodont Eutriconodonta is an order (biology), order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North America, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 ...

" mammal, recent studies have recovered it as a

triconodontid Triconodontidae is an extinct family of small, carnivorous mammals belonging to the order Eutriconodonta, endemic to what would become Asia, Europe, North America and probably also Africa and South America during the Jurassic through Cretaceo ...

eutriconodont Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the ...

, as most closely related to ''

Alticonodon ''Alticonodon'' is a genus of extinct mammal from the Late Cretaceous of North America. It is the geologically youngest known eutriconodont, and is a fairly more specialised animal than earlier representatives of this clade. Description ''Altic ...

'' and '' Corviconodon'' (these are in turn each other's sister taxa).Thomas Martin, Jesús Marugán-Lobón, Romain Vullo, Hugo Martín-Abad, Zhe-Xi Luo & Angela D. Buscalioni (2015). A Cretaceous eutriconodont and integument evolution in early mammals. Nature 526, 380–384.

Biology

Because of its abundance on

lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...

and

estuary An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...

deposits and particular association with

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

-rich areas, it has been suggested that ''Astroconodon'' was an aquatic

, an assertion reinforced by a perceived functional similarity between its molars and those of

cetaceans Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...

and

pinnipeds Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammal, marine mammals. They comprise the extant taxon, extant family (biology ...

. Other researchers like

Zofia Kielan-Jaworowska Zofia Kielan-Jaworowska (25 April 1925 – 13 March 2015) was a Polish paleobiologist. In the mid-1960s, she led a series of Polish-Mongolian paleontological expeditions to the Gobi Desert. She was the first woman to serve on the executive commit ...

, however, were not convinced, noting that eutriconodont dentition cannot be easily compared to

placental Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsup ...

dentation, and that they generally have a shearing function as opposed to the non- occluding, grasping teeth of marine mammals. The prevalence on aquatic deposits, per her own words, "may well be real, but its significance is not clear", as like many other species found in

fluvial In geography and geology, fluvial processes are associated with rivers and streams and the deposits and landforms created by them. When the stream or rivers are associated with glaciers, ice sheets, or ice caps, the term glaciofluvial or fluviog ...

deposits it may have simply been a terrestrial species whose remains were carried over by the waters, though she does cite Goldberg 2000 and recognises that there are stratifications of aquatic and terrestrial species to be noted. Terrestrial or aquatic, ''Astroconodon'', like most eutriconodonts, was almost certainly a predator. Like in most eutriconodonts, its triconodont molars were adapted for shearing, much ike the

carnassial Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...

s of

theria Theria (; Greek: , wild beast) is a subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes. Ch ...

n carnivores.Zofia Kielan-Jaworowska, Richard L. Cifelli, Zhe-Xi Luo (2004). "Chapter 7: Eutriconodontans". Mammals from the Age of Dinosaurs: origins, evolution, and structure. New York: Columbia University Press. pp. 216–248. .

Palaeoecology

''Astroconodon'' occurs in several fossil formations, most depicting

wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...

and

delta Delta commonly refers to: * Delta (letter) (Δ or δ), a letter of the Greek alphabet * River delta, at a river mouth * D ( NATO phonetic alphabet: "Delta") * Delta Air Lines, US * Delta variant of SARS-CoV-2 that causes COVID-19 Delta may also ...

environments. In both the

and the

it occurs alongside iconic

dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

such as ''

Tenontosaurus ''Tenontosaurus'' ( ; ) is a genus of medium- to large-sized ornithopod dinosaur. It was a relatively medium sized ornithopod, reaching in length and in body mass. It had an unusually long, broad tail, which like its back was stiffened with a n ...

'' and ''

Deinonychus ''Deinonychus'' ( ; ) is a genus of dromaeosaurid theropod dinosaur with one described species, ''Deinonychus antirrhopus''. This species, which could grow up to long, lived during the early Cretaceous Period, about 115–108 million ye ...

'', and in former it co-exists with a wide variety of other mammals including two other eutriconodonts, the considerably larger ''

Jugulator amplissimus ''Jugulator'' is an extinct genus of mammals from the Cretaceous of North America. It contains one species, ''Jugulator amplissimus''. A eutriconodont, it is known from the Cedar Mountain Formation, and is both a large sized and possibly ecolog ...

'' and closely related '' Corviconodon utahensis'', as well as many

multituberculate Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, a ...

s and

ns.

References

{{Taxonbar, from=Q12898868 Eutriconodonts Early Cretaceous mammals Mesozoic mammals of North America Cretaceous animals of North America Fossil taxa described in 1951 Taxa named by Bryan Patterson Prehistoric mammal genera